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Galapagos penguin (Spheniscus mendiculus), pair courting, Isabela Island, Galapagos Islands
Kingfisher ( Alcedo atthis) male and female perched on a root when building the nest, Lorraine, France, March
Spider wasp (Pompilidae) with spider prey, Buxa Tiger Reserve, India
Red fox (Vulpes vulpes) pair nuzzling. London, UK. February
Mistle thrush (Turdus viscivorus) calling perched on rowan tree. London, UK. December
Black-footed ferret (Mustela nigripes) running in the snow as it goes from one prairie dog burrow to another. Colorado, USA, Januay. January
Common swift (Apus apus) flying low to water while hunting, Norfolk, England, UK, July. Winner of Animal Behaviour Category of the British Wildlife Photography Awards 2019. Photographer
White-tailed sea eagle (Haliaeetus albicilla) flying over water with male Common eider prey (Somateria mollissima), Uto, Parainen, Finland, April
Male Black grouse (Tetrao / Lyrurus tetrix) in flight. Kuusamo, Finland, April
Boat-tailed grackle pair (Quiscalus major) in courtship display. Wakodahatchee Wetlands, Florida, USA, April
South African / Cape giraffe (Giraffa camelopardalis giraffa), two males necking / mock fighting, looking like a double headed animal, Zimanga Private Game Reserve, Zululand, South Africa
White-tailed eagle (Haliaeetus albicilla) flying over water with fish in talons. Norway. August
Great grey owl (Strix nebulosa) in flight in woodland during snowfall. Kuusamo. Finland. February
Apollo butterfly (Parnassius apollo) female laying eggs on branch. Parainen, Finland. July
Male Zander or Pikeperch (Stizostedion / Sander lucioperca) with mouth wide open, defending nest. Lake di Lugano or Ceresio, Ticino, Switzerland
Lowland copperhead snake (Austrelaps superbus) male basking on rusty trailer, on farm in evening. Melbourne, Victoria, Australia
Blotched blue-tongue skink (Tiliqua nigrolutea) female gaping with tongue out, in defense against perceived threat. Lake Tantangara, Snowy Mountains, New South Wales, Australia
Arctic fox (Vulpes lagopus), blue colour morph in winter coat walking through snow. Hornstrandir Nature Reserve, Iceland. February
Atlantic salmon (Salmo salar) leaping up waterfall to reach spawning grounds upstream. River Endrick, Loch Lomond and The Trossachs National Park, Scotland, UK. July
Lawess parotia (Parotia lawesii) male displaying. Papua New Guinea
Caribbean reef squid (Sepioteuthis sepioidea) eating fish at night, portrait. The Bahamas
Tiger cardinalfish (Cheilodipterus macrodon) male mouth brooding a clutch of eggs behind his sharp teeth. Gubal Island, Egypt. Strait Of Gubal, Red Sea
Pair of Rock beauty angelfish (Holacanthus tricolor) spawning at dusk. The larger male rubs against the flank of the female during the spawning rise
A school of Blackfin barracuda (Sphyraena qenie) forming the number 6 as they circle in deep water adjacent to a coral reef, Yolanda Reef, Ras Mohammed Marine Park, Sinai, Egypt. Red Sea. July
Common tern (Sterna hirundo) flying low over water. Hjalstaviken, Uppland, Sweden
Kottigehar dancing frog (Micrixalus kottigeharensis), male waving foot and calling, territorial behaviour. Agumbe, Western Ghats, India. Endemic
European otters (Lutra lutra) play fighting in the snow. Captive, Germany
Red Deer (Cervus elaphus) stags fighting during the rutting season. Bushy Park, London, UK. October
Giant barrel sponge (Xestospongia testudinaria) releases a large cloud of gametes as it spawns on a coral reef, while a Threadfin hawkfish (Cirrhitichthys aprinus) looks
Humpback whale (Megaptera novaeangliae) spouting, rainbow effect created in spray, aerial view. Baja California, Mexico. March
Pin-tailed whydah (Vidua macroura) male in display flight. Allahein River, Gambia
RF - African Lion cubs (Panthera leo) playing - age 4 months, Big Marsh, near Ndutu, Nogorongoro Conservation Area / Serengeti National Park, Tanzania
Red fox (Vulpes vulpes) cub in pasture asking vixen for food, England
Eurasian coot (Fulica atra) males fighting over territory. London, England, UK. December.;
Shoebill stork (Balaeniceps rex) eating a fish in the swamps of Mabamba, Lake Victoria, Uganda
Sockeye salmon (Oncorhynchus nerka) males fighting in river Adams River, British Columbia, Canada. October
Black Grouse (Lyrurus tetrix) male displaying with bokeh effect in the foreground, Liminka, Finland, March
Two Black woodpeckers (Dryocopus martius) males on tree trunk in territorial dispute, Peerdsbos, Braschaat, Belgium. April
Atlantic spotted dolphin (Stenella frontalis) juvenile, without spots, porposing, Tenerife, Canary Islands
Galapagos racer snake (Pseudalsophis biserialis) feeding on marine iguana hatchling, Cape Douglas, Fernandina Island, Galapagos
Racer snake (Pseudalsophis biserialis) feeding on lava lizard, Puerto Pajas, Isabela Island, Galapagos
Magnificent frigatebirds (Fregata magnificens) diving for fish prey, Borrero Bay, Santa Cruz Island, Galapagos
Magnificent frigatebird (Fregata magnificens) attacking Blue footed booby (Sula nebouxii) feeding chick. This bird is trying to steal the food from the parent. Santa Cruz Island, Galapagos
Espanola saddelback tortoise (Chelonoidis hoodensis) pair mating, Espanola Island, Galapagos
Blue-footed booby (Sula nebouxii) with chicks begging for food, Santa Cruz Island, Galapagos, Ecuador
Seven-spot ladybird (Coccinnella septempunctata) just emerged from hibernation, sunning on a Honeysuckle leaf with buildings in the background, Wiltshire garden, UK, April
Red-winged Blackbird (Agelaius phoenicus) male singing from snow-covered red maple (Acer rubrum) in flower, Ithaca, New York, USA. April 2020
Wild Turkey (Meleagris gallopavo) male displaying in early spring, in light snow, New York, USA