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Lions (Panthera leo) five drinking together, Ndutu, Tanzania
Lionesses (Panthera leo) grooming each other, Masai-Mara Game Reserve, Kenya
Bornean orangutan (Pongo pygmaeus) mother and babies, Tanjung Puting National Park, Borneo-Kalimatan, Indonesia, endangered species
Jaguar (Panthera onca) mother playing with four month old cub, native to Southern and Central America, captive
Erect-crested penguin (Eudyptes sclateri) pair in greeting display. Antipodes Island, New Zealand Sub-Antarctic Islands. Endemic to Antipodes and Bounty Islands. Endangered species
Lionesses (Panthera leo) trying to bring down African buffalo (Syncerus caffer) Londolozi Private Game Reserve, Sabi Sand Game Reserve, South Africa. Sequence 5 of 5
RF - Leopard (Panthera pardus) cub playing with mothers tail, Londolozi Private Game Reserve, Sabi Sand Game Reserve, South Africa
RF - Leopard (Panthera pardus) stalking prey, Londolozi Private Game Reserve, Sabi Sand Game Reserve, South Africa
Arctic fox (Vulpes lagopus) in summer coat calling, Wrangel Island, Far Eastern Russia, August
Arctic fox (Alopex lagopus) juveniles play fighting, Wrangel Island, Far East Russia, August
Leopard (Panthera pardus) adult with scar accorss face licking paw, Londolozi Private Game Reserve, Sabi Sand Game Reserve, South Africa
Snow goose (Chen caerulescens caerulescens) brooding chicks, with rusty orange face from iron rich soil in which it forages. Wrangel Island, Far Eastern Russia, June
Emperor penguins (Aptenodytes forsteri) blizzard near Snow Hill Island colony in Weddell Sea, Antarctica
Dipper (Cinclus cinclus) displaying or drying wings, Kuusamo Finland March
RF- Icelandic horse showing flehmen response, Iceland. July
A group of Haflinger, Pure Arab, Shagya Arab and East Bulgarian fillies and mares running in snow, Kabiuk National Stud, Shumen, Bulgaria
Wild Konik horses with Starlings (Sturnus vulgaris) Odry delta reserve, Stepnica, Poland, July
An Einsiedler / Swiss warmblood mare (Equus caballus) cantering, Schwyz, Switzerland, July
Kottigehar dancing frog (Micrixalus kottigeharensis) male calling and displaying to attract female by waving foot, Agumbe, Western Ghats, India
Asiatic elephants (Elephas maximus), dust bathing at dawn. Jim Corbett National Park, India
Kottigehara dancing frog (Micrixalus kottigeharensis), dancing frog name given because of foot flagging behavior between competing males. Endemic to Western Ghats. Agumbe, Karnataka, India
Anils bush frog (Raorchestes anili) inflating vocal sac to attract mate, Endemic to Western Ghats. Coorg, Karnataka, India
Jayarami Bush frog (Raorchestes jayarami), inflating vocal sac, calling to attract mate, Endemic to Western Ghats. Anamalai Wildlife Sanctuary, India
Humayuns night frog (nyctibatrachus humayuni), male guarding cluster of eggs on leaf. Western Ghats Amboli, Maharashtra, India. Endemic
Low angle view of Demoiselle crane (Anthropoides virgo) flying and feeding during winter migration. Khichan, Western Rajasthan, India. December
Demoiselle crane (Anthropoides virgo) low angle view of birds flying and landing near the chugga ghar (bird feeding enclosure ) in Khichan Village, durin their annual migration
Indo-Pacific humpback dolphins (Sousa chinensis) surfacing, Tai O, Lantau Island, Hong Kong, China
Common green magpie (Cissa chinensis) bathing, Hong Bung He, Dehong, Yunnan, China
Minke whale ( Balaenoptera acutorostrata) breaching, Bay of Fundy, Canada, July
Leaf mantis (Choeradodis sp. ) Tortuguero National Park, Costa RicaLeaf mantis (Choeradodis sp.) Tortuguero National Park, Costa Rica
Long beaked common dolphin (Delphinus capensis) pod porpoising Baja California, Mexico
Firecrest (Regulus ignicapilla) singing, Wiltshire, England, UK, April
Ants (Formicidae) attacking newly emerged cicada (Cicadacae), Yasuni National Park, Ecuador. Highly commended in the Invertebrates Category of the Wildlife Photographer of the Year Awards (WPOY)
Sedge Warbler (Acrocephalus schoenobaenus) Greylake RSPB Reserve, Somerset Levels, England, UK, August
Atlantic salmon (Salmo salar) jumping up waterfall during spawning migration upstream, Ume River, Sweden. July. Photographed for The Freshwater Project
Flightless cormorant (Phalacrocorax harrisi) pair in courtship Puerto Pajas, Isabela Island, Galapagos
Split level view of Brown pelicans (Pelecanus occidentalis) feeding, Tortuga Bay, Santa Cruz Island, Galapagos
Brown pelican (Pelecanus occidentalis) feeding chicks at nest, Puerto Ayora / Academy Bay, Santa Cruz Island, Galapagos
Spoonbill (Platalea leucorodia) carrying nest material, The Netherlands
Kerry Hill domestic sheep, spring lambs running, England, UK
Atlantic yellow-nosed albatross (Thalassarche chlororhynchos) courtship display, Gough Island, Gough and Inaccessible Islands UNESCO World Heritage Site, South Atlantic
Gough Island bunting (Rowettia goughensis) gathering nesting material. Gough Island, Gough and Inaccessible Islands UNESCO World Heritage Site, South Atlantic. Endemic. Critically endangered species
European barracuda (Sphyraena sphyraena) and Bluefish (Pomatomus saltatrix) circling baitball of Atlantic horse mackerel (Trachurus trachurus) Formigas Islets, Azores
Rabbit grooming toes, Okunoshima Rabbit Island, Takehara, Hiroshima, Japan
Feral domestic rabbit (Oryctolagus cuniculus) males fighting one leaping into the air, Okunojima Island, also known as Rabbit Island, Hiroshima, Japan
American oystercatcher (Haematopus palliatus) feeding on Calico Clam, shortly after dawn, seashore, Tampa Bay, St. Petersburg, Florida, USA. April
Whitebelly damselfish (Amblyglyphidodon leucogaster) pair spawning / laying eggs on a seafan (Annella mollis). Ras Mohammed National Park, Sinai, Egypt. Red Sea
Cheetah (Acinonyx jubatus), cubs age 8 weeks playing, Masai Mara Game Reserve, Kenya
