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Barn swallow (Hirundo rustica) alighting at pond to collect mud for nest building, Scotland, UK, May
Oceanic whitetip shark (Carcharhinus longimanus) accompanied by pilotfish (Naucrates ductor) as it cruises beneath the surface of the Red Sea in the late afternoon, close to Little Brother Island
Common goldeneye (Bucephala clangula) male performing its mating display. Southern Norway. March
Young ground squirrels (Xerus inauris) playing, Kgalagadi Transfrontier Park, South Africa. Non-ex
Group of California sea lions (Zalophus californianus) swimming in kelp forest (Macrocystis pyrifera), California, USA. Pacific ocean
Egyptian cobra (Naja haje) with head raised up and hood expanded, near Ouarzarte, Morocco
Black Cocker Spaniel puppy, against white background
Feral domestic rabbit (Oryctolagus cuniculus) cleaning its face, Okunojima Island, also known as Rabbit Island, Hiroshima, Japan
Golden Retriever dog pup, Oscar, 3 months, in play-bow, against white background
Domestic dog, Dogo Argentino, three puppies playing with stick, France
Prairie Rattlesnake (Crotalus viridis) sunbathing, Bozeman, Montana, USA
Rufous-tailed hummingbird (Amazilia tzacatl) territorial fighting around Heliconia flower (Heliconia latispatha) La Selva, Costa Rica
Tree swallows (Tachycineta bicolor), perched on wire, calling aggressively to each other, Madison River, Montana, USA, June
Black flat-coated retriever dog splashing in water and retrieving stick (Canis familiaris)
Two Irish Gypsy cob mares (Equus caballus), one dapple grey - grazing and one piebald - walking on rough pastureland on Hackpen Hill with a Beech clump (Fagus sylvaticus) in the background
Geranium Argus butterfly (Plebejus eumedon) feeding on Wood Cranesbill (Geranium sylvaticum) northern Finland, June
Humpback Whale (Megaptera novaeangliae) tail slapping, tail lob, lobtailing. Vancouver Island, British Columbia, Canada, July
Male Ostrich (Struthio camelus) protecting chicks from the sun with its wings, Samburu National Reserve, Kenya, Africa
White-tailed eagle (Haliaeetus albicilla) in flight, Norway, April
Indri (Indri indri) calling, Andasibe-Mantadia NP, Madagascar
Lion(Panthera leo) cub playing with its mothers tail, Masai-Mara Game Reserve, Kenya. Vulnerable species
Brush footed butterfly (Lycorea sp. ) caterpillar with parasitic wasp cocoons on silk threadsBrush footed butterfly (Lycorea sp.) caterpillar with parasitic wasp cocoons on silk threads, Yasuni National Park, Orellana, Ecuador
Juvenile of Amazon Tree-Boa (Corallus hortulanus) in defensive position. Yasuni National Park, Orellana, Ecuador
Grey-cheeked parakeets (Brotogeris pyrrhoptera) perched and grooming on a branch. Guayaquil, Guayas, Ecuador
Indian peafowl (Pavo cristatus) peacock walking and displaying feathers, Yala National Park, Southern Province, Sri Lanka
Kinabalu giant red leech (Mimobdella buettikoferi) feeding on Kinabalu giant earthworm (Pheretima darnleiensis), on Mount Kinabalu, Borneo
Raja Brookes Birdwing Butterfly (Trogonoptera brookiana), Borneo
Mountain tree shrew (Tupaia montana) feeding on nectar secreted by the endemic Pitcher Plant (Nepenthes kinabaluensis) Montane forests (at 2200m-3000m), slopes of Mt Kinabalu
Great Fen / Raft spider (Dolomedes plantarius), adult female eating an invasive species of fish, Western mosquitofish, (Gambusia affinis), Alessandria, Italy, August
Cat gecko (Aeluroscalabotes felinus) with coiled tail, Sarawak, Malaysian Borneo
Wavy-lined emerald moth caterpillar (Synchlora aerata) which camouflages itself by attaching parts of flower to itself, on Black-eyed susan (Rudbeckia hirta) Morris Arboretum, Philadelphia
Two Jaguars (Panthera onca) snarling at each other on the riverbank, Pantanal, Brazil. September
Northern Hawk Owl (Surnia ulula) preening, Varanger, Norway. June
Spotted owlet (Athene brama) pair on a branch, Bharatpur / Keoladeo Ghana National Park, India
RF- Indian eagle owl (Bubo bengalensis) head portrait, captive, from India
Mountain hare (Lepus timidus) grooming itself, with back foot raised, Cairngorms National Park, Scotland, UK, February. Highly Commended in the Animal Behaviour category of the British Wildlife
RF - Female emperor dragonfly (Anax imperator) laying eggs on garden pond, Broxwater, Cornwall, UK. July
Adder (Vipera berus) basking amongst bracken fronds. Peak District National Park, Derbyshire, UK. April
Otter (Lutra lutra) female rolling around in seaweed, Mull, Scotland, England, UK, September
Red deer (Cervus elaphus) stag bellowing during rut, at night with full moon, Derbyshire, UK
Otter (Lutra lutra) female grooming in seaweed, Mull, Scotland, England, UK, September
Hazel Dormouse (Muscardinus avellanarius) hibernating amongst leaves and acorns. Captive. Leicestershire, UK, October
Raven (Corvus corax) adult calling from rock in mountain habitat, Scotland, UK. November. Third place in British Birds Bird Photograph of the Year competition 2016
Flock of Common Starling (Sturnus vulgaris) gathering before landing at winter roost, Salisbury Plain, Wiltshire, UK, January
Gannet (Sula bassana) diving, July UK Shetland
Adult male Sparrowhawk (Accipiter nisus) with prey perched on a branch, Dumfries, Scotland, UK, November
White spotted eagle ray (Aetobatus narinari) feeding by digging in the sand and seagrass for invertebrates. East End, Grand Cayman, Cayman Islands, Caribbean Sea
The vertical reef wall at Shark Reef, Ras Mohammed, with Scalefin anthias (Pseudanthias squamipinnis) and soft corals (Dendronephthya spp.) Yellowsaddle goatfish (Parupeneus cyclostomus)