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Hooded nudibranchs (Melibe leonina) filter feeding beneath bull kelp. Hussar Point, Browning Pass, Port Hardy, Vancouver Island, British Columbia, Canada
RF- School of Blackfin barracuda (Sphyraena qenie) forming circle in open water at Shark Reef, Ras Mohammed Marine Park, Sinai, Egypt. Red Sea
Male Yellow-headed jawfish (Opistognathus aurifrons) blows out the clutch of eggs he is incubating in his mouth to oxygenate them. East End, Grand Cayman, Cayman Islands, British West Indies
A female (in front) and male Sockeye salmon (Oncorhynchus nerka) over eggs in their spawning river. Adams River, British Columbia, Canada, October
Pair of Sockeye salmon (Oncorhynchus nerka) on their redd in a shallow stream. Female in front of male. Huihil Creek, Adams River, British Colombia, Canada, October
Striped mackerel (Rastrelliger kanagurta) mouth wide open as it swims through the water, filtering zoopankton with its gill rakers. Marsa Shouna, Port Ghalib, Marsa Alam, Egypt. Red Sea
Diver swimming with an aggregation of Cod (Gadus morhua) over a kelp forest. These cod were gathered in early spring off the north coast of Iceland to spawn. Thorshofn, Iceland. North Atlantic Ocean
Guianan Cock-of-the-rock (Rupicola rupicola), during courtship display, Guianan Shield, Suriname
Oman anemonefish / clownfish (Amphiprion omanensis) in a a Bulb-tentacle sea anemone (Entacmaea quadricolor), coast of Dhofar and Hallaniyat islands, Oman. Arabian Sea
Mountain tree shrew (Tupaia montana) feeding on nectar secreted by the endemic Pitcher Plant (Nepenthes kinabaluensis) Montane forests (at 2200m-3000m), slopes of Mt Kinabalu
Leopard (Panthera pardus) hunting Springbok (Antidorcas marsupialis) Etosha, Namibia, Finalist in the Mammals Category of the Wildlife Photographer of the Year 2015
Hyacinth macaws (Anodorhynchus hyacinthinus) mutual preening, Estrada Parque, Pantanal, Brazil
Great frigatebird (Fregata minor) male displaying inflated gular sac, Galapagos, Ecuador
Lion (Panthera leo) cubs resting, Masai-Mara Game Reserve, Kenya
RF - Sea otter (Enhydra lutris) floating, California, USA, July
Black-and-white Jack-a-poo, Jack Russell cross Poodle puppy with ears pointing in opposite directions
Burrowing owl (Athene cunicularia) fledgling with head on side, Pantanal, Brazil. November
Arctic fox (Vulpes lagopus) with Snow goose egg in mouth, mid moult from winter to summer fur, Wrangel Island, Far Eastern Russia, May
Great white egret (Ardea alba) splashing, Hungary, January
Barn Owl (Tyto alba) hunting / hovering, Somerset, UK, trained birdBarn Owl (Tyto alba) hunting/hovering, Somerset, UK, trained bird
Emma Grays Forest Lizard (Calotes emma) in threat display, Xishuangbanna National Nature Reserve, Yunnan Province, China, March
RF - California sea lion (Zalophus californianus) with mouth open close to the camera. The milk teeth of this pup show decay, which is quite usual. Los Islotes, La Paz, Baja California Sur, Mexico
California sea lion (Zalophus californianus) uses a Panamic cushion star (Pentaceraster cumingi) as a toy. The sealions pick up the starfish and then drop them and chase after them as they sink
Mandarinfish (Synchiropus splendidus) pair spawning. The larger male (behind) is lifting the female into the spawning rise on his pectoral fin, the smaller female (foreground) is releasing eggs
Blue spotted trevally (Caranx bucculentus) hunting small fish in the reef, South Ari Atoll, Maldives, Indian Ocean
Red Sea anemonefish (Amphiprion bicinctus) in their home, a Sea anemone (Heteractis magnifica), on a coral reef. Big Brother island, Red Sea. Egypt
Green sea turtle female (Chelonia mydas) returning to the sea, Tortuguero National Park, Costa Rica
RF - Scarlet macaw (Ara macao) pair preening, Osa Peninsula, Costa Rica
Green-winged macaws (Ara chloroptera) preening each other. Brazil. South America
Reticulated giraffe (Giraffa camelopardalis reticulata) mother grooming baby, Samburu Game Reserve, Kenya, Africa, August
Toco Toucan (Ramphastos toco) beak open with tongue visible while feeding on mango, Brazil
Atlantic Puffin (Fratercula arctica) in flight, coming into land with beak full of sand eels (Ammodytes tobianus). Fair Isle, Shetland Islands, Scotland, UK, July
Black howler monkeys (Alouatta caraya) male and two females calling from tree, captive, Apenheul Park, Netherlands
Polar bear (Ursus maritimus) juvenile rolling around on newly formed pack ice, Beaufort Sea, off the 1002 area of the Arctic National Wildlife Refuge, North Slope, Alaska
Loggerhead sea turtle (Caretta caretta) emerging from shell, Dalyan delta, Turkey, July
Lion (Panthera leo) cub portrait, Masai Mara Game Reserve, Kenya
Indian fox (Vulpes bengalensis) pups at play by a den in the Little Rann, Kutch, Gujarat, India. Honorable mention in the Land Mammals category of the BigPicture Competition 2014
Grevy zebra (Equus grevyi) stallions fighting, Samburu National Reserve, Kenya
Three False clownfish (Amphiprion ocellaris) in Sea anemone (Heteractis magnifica) Lighthouse Reef, Cabilao Island, Bohol, Central Visayas, Philippines, Pacific Ocean
Atlantic salmon (Salmo salar) migrating for spawning in river, Gaspe Peninsula, Quebec, Canada, October
Arctic tern (Sterna paradisaea) in flight with fish in beak for feeding chick, Machias Seal Island, Bay of Fundy, New Brunswick, Canada, July
Tropical shield mantis (Choeradodis rhombicollis) in defensive position, on flower, Chinambi, Carchi, Ecuador
Brown throated three toed sloth (Bradypus variegatus) hanging from branch, Yasuni National Park, Orellana, Ecuador
African elephants (Loxodonta africana) swimming across Chobe River, trunks raised above surface for breathing, Chobe National Park, border between Botswana and Namibia
Pod of Risso Dolphin (Grampus griseus) at surface, Baja California, Mexico
Large pod of Bottlenose dolphins (Tursiops truncatus) porpoising over waves during annual sardine run, Port St Johns, South Africa
Humpback whale (Megaptera novaeangliae australis) courtship, with the female above showing her abdomen and the male below. Vava'u, Tonga. Pacific Ocean
Humpback whale (Megaptera novaeangliae), fluke of female with male calf playing below, Vava'u, Tonga, Pacific Ocean
