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Andean cock-of-the-rock (Rupicola peruvianus), two males perched in tree at lek in mid-altitude montane rainforest. Manu Biosphere Reserve, Amazonia, Peru
Parson's chameleon (Calumma parsonii), male hunting in forest canopy. Mitsinjo Forest, Andasibe-Mantadia National Park, eastern Madagascar
Bichon frise beside Colorado blue columbine (Aquilegia coerulea) flowers. Old Lyme, Connecticut, USA. May
Bichon frise standing on rock amongst Rhododendron flowers. Haddam, Connecticut, USA. May
RF - Andean cock-of-the-rock (Rupicola peruvianus), male perched in tree at a lek. Mid-altitude montane rainforest, Manu Biosphere Reserve, Amazonia, Peru
Blackbird (Turdus merula), male singing whilst perched in Ash (Fraxinus excelsior) tree, near Bath, England, UK. May
Grey whale (Eschrichtius robustus) breaching, San Ignacio Lagoon, El Vizcaino Biosphere Reserve, Baja California, Mexico, March
RF - Playful Silver tabby kitten rolling on back
White tailed sea eagle (Haliaeetus albicilla) perched on tree stump with fish. Hooded crow (Corvus cornix) flying in to attack in background. Danube Delta, Romania. May
Bee-eater (Merops apiaster) with Bumblebee prey, Sierra de Grazalema Natural Park, southern Spain, July
Whip scorpion (Heterophrynus elephas) hunting for food on a large tree root of the rainforest. Los Amigos Biological Station, Peru
Stag beetle (Lucanus cervus) males fighting on oak tree branch, Elbe, Germany, June
Black spider monkey (Ateles chamek) calling, captive, Peru
Indo-Pacific humpback dolphin (Sousa chinensis) with fishing net damage around throat, Pearl River Estuary, Lantau Island, Hong Kong, China
RF - White lion (Panthera leo) male, portrait of head. Captive, Netherlands
RF - Meerkat (Suricata suricatta) resting on cool sand, Kgalagadi Transfrontier Park, South Africa
Straw-coloured fruit bat (Eidolon helvum), female flying carrying pup on front. Lamin, Gambia
Yacare caiman (Caiman yacare) with butterfly on snout, Cuiaba River, Pantanal Matogrossense National Park, Pantanal, Brazil
Yacare caiman (Caiman yacare) on river bank, Cuiaba River, Pantanal Matogrossense National Park, Pantanal, Brazil
Jaguar (Panthera onca) male, hunting Capybara (Hydrochoerus hydrochaeris). The capybara while jumping away has cut the nose of the Jaguar with its toemail
Jaguar (Panthera onca) male, chasing a Caiman. Cuiaba River, Pantanal Matogrossense National Park, Pantanal, Brazil
African lion (Panthera leo) growling at potential danger in the water (Panthera leo) Okavango Delta, Botswana
Lioness (Panthera leo) resting up a tree - only three populations of lions are known to do this habitually, Ishasha Sector, Queen Elizabeth NP, Uganda
Lion (Panthera leo) cub up a tree - only three populations of lions are known to do this habitually, Ishasha Sector, Queen Elizabeth NP, Uganda
African lion (Panthera leo) three subadults resting on the road, one snarling, Central Kalahari Game Reserve, Botswana
Jaguar (Panthera onca) mother grooming with four month old cub, native to Southern and Central America, captive
RF- Jaguar head profle portrait (Panthera onca) head portrait captive. Pantanal, Brazil
Mountain gorillas (Gorilla gorilla beringei) grooming, members of the Munyaga group, Virunga National Park, North Kivu, Democratic Republic of Congo, Africa, Critically endangered
Polar bear (Ursus maritimus) sow jumping while hunting for seals on sea ice, off the coast of Svalbard, Norway
Tabby cats, Picasso and Smudge, age 3 months, together. Smudge with his paws up on Picasso
Northern goshawk (Accipiter gentilis) perched on snag with grouse prey, Finland, March. March
Western capercaillie (Tetrao urogallus) Tver, Russia. May
Black Grouse (Tetrao tetrix) displaying at a lek in snow, Tver, Russia. April
Arctic fox (Vulpes lagopus) with Snow goose egg in mouth, mid moult from winter to summer fur, Wrangel Island, Far Eastern Russia, June
RF - Polar Bears (Ursus maritimus) males fighting, Churchill, Canada, November
Camargue horse, two, one with mouth open. Bouches du Rhone, France May
Two wild Mustang stallions fighting in the White Mountain Herd Area, Wyoming, USA. August
Wild Mustang horse rearing, Pryor mountains, Montana, USA. June
Icelandic horse with mouth open, Iceland
Irish Gypsy cob (Equus caballus) grey stallion and piebald mare grazing on rough pastureland on Hackpen Hill with a Beech clump (Fagus sylvaticus) in the background, The Ridgeway
Grey Irish Gypsy cob stallion (Equus caballus) grazing rough pastureland on Hackpen Hill with Oilseed rape (Brassica napus) and other arable crops in the background, The Ridgeway
Chinese cobra (Naja atra) in threat stance, Shek Pik, southwestern coast of Lantau Island, Hong Kong, China
Indo-Pacific humpback dolphin ( Sousa chinensis) surfacing, Tai O, Lantau Island, Hong Kong, China
Atlantic salmon (Salmo salar) portrait, migrating to spawn in river, Gaspe Peninsula, Quebec, Canada, October
Mangrove hummingbird (Amazilia boucardi) female feeding chicks in nest, Pacific coast mangroves area, Costa Rica, Endangered species
Northern gannet (Morus bassanus) pair, courtship display, Machias Seal Island, Bay of Fundy, New Brunswick, Canada, May
Minke whale (Baelanoptera acutorostrata) breaching, Bay of Fundy, New Brunswick, Canada, July
Harbour porpoise, (Phocoena phocoena) porpoising, Bay of Fundy, New Brunswick, Canada, August