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Wood pigeon (Columba palumbus) flying with nesting material in beak, Lorraine, France, April
Timber rattlesnake (Crotalus horridus) female with newborn young, Pennsylvania, USA. September
Great grey owl (Strix nebulosa) hunting over snow, woodland in background. Kuusamo, Finland. February
Great grey owl (Strix nebulosa) close up hunting in snow. Kuhmo, Finland. February
Elk (Cervus canadensis) bull and cw sniffing noses, Yellowstone National Park, Wyoming, USA. September
Rocky mountain elk (Cervus elaphus nelsoni) bull bugling during rut. Rocky Mountain National Park, Colorado, USA. October
Sahara sand viper (Cerastes vipera) burrowing into sand to hide, captive, occurrs Mauritania to Egypt, Africa. Sequence 5 of 5
Salmon shark (Lamna ditropis). Port Fidalgo, Alaska, USA. July
Leopard (Panthera pardus) cubs age four months playing, Jao Reserve, Okavango, Botswana
Osprey (Pandion haliaetus) carrying fish above river in Taipei, Taiwan
Dugong (Dugong dugon) male and juvenile Golden trevally (Gnathanodon speciosus) shoal swimming over Turtlegrass (Halophila stipulacea) seagrass bed. Marsa Alam, Egypt
Common collared lizard (Crotaphytus collaris auriceps) male basking on rock. Arches National Park, Utah, USA. May
North American Beaver (Castor canadensis) grooming. Acadia National Park, Maine, USA
African fish eagle (Haliaeetus vocifer) attacking a female Saddle-billed Stork (Ephippiorhyncus senegalensis )trying to steal a fish it has just caught. Liuwa Plains National Park, Zambia
RF - Tabby kitten dancing. (This image may be licensed either as rights managed or royalty free.)
Andalusian horsea, two stallions coming together in arena one dappled grey and one pure white. Spain
Iberian lynx (Lynx pardinus) walking on frosty grass, Parque Natural Sierra de Andujar, Andalucia, Spain. January
Rabbit (Oryctolagus cunniculus) juvenile grooming ears, Wiltshire, UK, May. Sequence 1 of 3
Yellowfin tuna (Thunnus albacares) hunting mackerel (Scomber colias) Tenerife, Canary Islands
Portrait of a wild adult female Bobcat (Lynx rufus) with Hispid cotton rat (Sigmodon hispidus) prey, Texas, USA. September
Portrait of a wild female Bobcat (Lynx rufus) kitten playing, Texas, USA. September
Tokay gecko (Gekko gecko) enacting a defensive display towards a perceived threat, Phuket Island, Thailand, March. Controlled conditions
Magnificent riflebird (Ptiloris magnificus alberti) male, on his display perch performing a display. Piccaninny Plains Sanctuary, Cape York Peninsula, Queensland, Australia
RF-Asian elephant (Elephas maximus) dust bathing. Jim Corbett National Park, Uttarakhand, India. (This image may be licensed either as rights managed or royalty free)
Lion (Panthera leo) males mock fighting / play fighting. Ngorongoro Crater, Ngorongoro Conservation Area (NCA), Tanzania
Southern right whale (Eubalaena australis) diving, with tail fluke splash. Monumento Natural Ballena Franca Austral, UNESCO World Heritage Site, Valdes Peninsula, Patagonia, Argentina
African lion (Panthera leo) and lioness after mating, Masai Mara National Reserve, Kenya
Lady Amhersts pheasant (Chrysolophus amherstiae) male displaying, Kanding, Sichuan, China. April
European rabbits (Oryctolagus cuniculus) fighting each other, Kiskunsag National Park, Hungary. June. Winner, 2019 Cadiz Photo Nature Competition
Polar bears (Ursus maritimus) fighting in water. Beaufort Sea, Kaktovik, Alaska, USA. Highly commended in the Polar Passion category of the Natures Best Photography Competition 2019
Bee-eater (Merops apiaster) male offering wasp as nuptial gift, Hungary. June
Timber rattlesnake (Crotalus horridus) females and newborn young at maternity site. Pennsylvania, USA. August
Fosa or Fossa (Cryptoprocta ferox)(sometimes erroneously fossa) male in dry deciduous forest. Kirindy, western Madagascar
Guanacos (Lama guanicoe), two males fighting, trying to over power opponent and bite each others testicles. Torres del Paine National Park, Patagonia, Chile. December
Bush cricket / Katydid (Tettigoniidae ), female leaf mimic ovipositing into branch. Manu Biosphere Reserve, Peru
RF- Wild male Jaguar (Panthera onca palustris) running through shallows of backwater of Cuiaba River in late afternoon sunlight. Northern Pantanal, Brazil
Giant River Otter (Pteronura brasiliensis) feeding on Striped Catfish or Cachara (Pseudoplatystoma fasciatum) amongst water hyacinth. Tributary off the Cuiaba River, Mato Grosso, Pantanal, Brazil
North Pacific humpback whale (Megaptera novaeangliae kuzira) breaching at sunset, Kupreanof Island, Frederick Sound, Alaska, USA, July
Humpback whales (Megaptera novaeangliae) engaged in courtship, with the male hovering above the female. Vava'u, Tonga, Pacific Ocean
Twinspot snapper (Lutjanus bohar) shoal rushing up to spawn, releasing a cloud of milky white sperm and eggs in the water column. Caroline Islands, Palau, November
Ocean sunfish (Mola mola) and Pilot fish (Naucrates sp). Tenerife, Canary Islands
Bluestreak cleaner wrasse (Labroides dimidiatus) cleans among the sharp teeth of a predatory roving coral grouper (Plectropomus pessuliferus) beneath an overhang on a coral reef
Ruff (Philomachus pugnax) males displaying and fighting each other at a lek, Finland. May
Red fox (Vulpes vulpes) in mid air, snow diving / pouncing whilst hunting for rodents. Hayden Valley, Yellowstone National Park, USA. February
Dugong (Dugong dugon) male and juvenile Golden trevally (Gnathanodon speciosus) swimming over Seagrass (Halophila stipulacea) meadow. Marsa Nabaa, Marsa Alam, Egypt
Little red flying fox (Pteropus scapulatus) female and suckling baby in flight, searching for place to roost. Nitmiluk National Park, Northern Territory, Australia
Cape buffalo (Syncerus caffer caffer) portrait, Red-billed oxpeckers (Buphagus erythrorhynchus) removing parasites. Nakuru, Kenya
Turtle dove (Streptopelia turtur) female feeding chicks on nest. Ranmore Common, near Dorking Surrey, England, UK. July
