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Two Red Foxes (Vulpes vulpes) fighting. The Netherlands. August
Great Egrets (Ardea alba) territorial fight above nest colony, Everglades National Park, Florida, USA, February
Ruffs displaying and fighting (Philomachus pugnax) Agapa River, Taimyr Peninsula, Siberia, Russia
Arctic tern (Sterna paradisaea) adult hovering directly in front of the sun, defending nest from perceived threat, Farne Islands, UK. June Non-ex
Black Rhinoceros (Diceros bicornis) charging. Etosha National Park, Namibia, January
Two male Mute swans (Cygnus olor) displaying on water in territorial dispute, UK
Western Diamondback Rattlesnake (Crotalus atrox), adult in striking pose in wildflower field. Laredo, Webb County, South Texas, USA, April
Two Brown bears (Ursus arctos) fighting in woodland wetlands, Kuhmo, Finland, July
Feral domestic horses (Equus caballus) wild herd stallion (grey) fighting a bachelor chestnut stallion, Blue Swallow Reserve, near Kaapsehoop, Mpumalanga province / Eastern Transvaal, South Africa
European hare (Lepus europaeus) mating pair boxing in field, Slovakia
Arctic tern (Sterna paradisaea) in flight, vocalising, aggresive behaviour, Farne Islands, Northumberland, UK, June
Arctic tern (Sterna paradisaea) in flight, calling, Farne Islands, Northumberland, UK, June
A wild Misaki-uma breeding stallion (right) brings back one of his mares to his band while another breeding stallion (left) charges him away, in the Cape Toi Reserve, Miyazaki Prefecture
Eastern green mamba {Dendroaspis angusticeps} strike pose, captive, from East Africa
Mute swan (Cygnus olor) territorial adult, aggressive display posture, Derbyshire, UK, March
Arctic tern (Sterna paradisaea) adult aggressively defending its nest from a perceived threat, Farne Islands, UK
Male Red hartebeest (Alcelaphus buselaphus) fighting, Kgalagadi Transfrontier Park, Kalahari desert, South Africa
White rhinoceros {Ceratotherium simium} head on, Etosha national park, Namibia
Silhouette of Eastern Grey Kangaroo {Macropus giganteus} males boxing, New South Wales, Australia
Humpback whale lobtailing {Megaptera novaengliae} Dominican Republic, Caribbean
Two male Thomsons gazelles (Eudorcas thomsonii) fighting, Masai-Mara game reserve, Kenya
Confrontation between a great egret (Ardea alba) and grey heron (Ardea cinerea), Lake Csaj, Kiskunsagi National Park, Hungary
Eurasian coot (Fulica atra) males fighting over territory. London, England, UK. December.;