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Two male Dusky groupers (Epinephelus marginatus) fighting for territory during mating season at baixa do meros, a small pinnacle at Formigas islets Protected area, near Santa Maria Island, Azores
White tailed eagles (Haliaeetus albicilla) two juveniles fighting in flight, Lake Csaj, Kiskunsagi National Park, Pusztaszer, Hungary. January
Great egret (Ardea alba) fighting over fish with Grey heron (Ardea cinerea) Lake Csaj, Kiskunsagi National Park, Pusztaszer, Hungary. January
Two wild Mustang stallions fighting in White Mountain Herd Area, Wyoming, USA. August
White rhinoceros (Ceratotherium simum) two males fighting, in evening light, Sabi Sand Game Reserve, South Africa
Ruff (Philomachus pugnax) males fighting at lek, Varanger, Norway, May 2016
Common reed bunting (Emberiza schoeniclus) attacking Common cuckoo (Cuculus canorus), female, Braunschweig, Lower Saxony, Germany, May
Whooper swans (Cygnus cygnus) fighting, Finland, April
Cormorant (Phalacrocorax carbo) and Great egret (Ardea alba) fighting over fish, Lake Csaj, Kiskunsagi National Park, Pusztaszer, Hungary, January
King cobra (Ophiophagus hannah) juvenile in threat pose on white background, captive occurs in South Asia. Venomous species
Hippopotamus (Hippopotamus amphibius) bull emerging from water to charge, iSimangaliso Wetland Park, KwaZulu Natal National Park. South Africa
Sperm whale (Physeter macrocephalus) young male trying to copulate with a female. The calf of the female was in the way so the male is trying to chase it away. Dominica
San Cristobal giant tortoises (Chelonoidis chatamensis), fighting, Galapaguera, San Cristobal Island, Galapagos
Spotted nutcrackers (Nucifraga caryocatactes) fighting in snow, Vitosha Mountain, Sofia, Bulgaria
Puffin (Fratercula arctica) with fish chased by Lesser black-backed gull (Larus fuscus) Isle of May, Firth of Forth, Fife, Scotland, UK, June
Two wild Namib stallions fighting, Namib Nakluft National Park, Namib Desert, Namibia, September 2008
White-rumped snowfinch (Montifringilla taczanowskii) two fighting, Tibetan Plateau, Qinghai, China
RF - Red deer (Cervus elaphus) stags fighting, Holkham Park, Norfolk, England, UK, September
Raggy scorpionfish (Scorpaenopsis venosa) disputing territory. Ambon Bay, Ambon, Maluku Archipelago, Indonesia. Banda Sea, tropical west Pacific Ocean
Northerns (Accipiter gentilis) fighting over squirrel carcass, Finland, March MarchNortherns (Accipiter gentilis) fighting over squirrel carcass, Finland, March.. March
Black Grouse (Tetrao tetrix) males fighting in winter, Tver, Russia. April
Northern gannets (Morus bassanus) on rock squabbling, Saltee Islands, Ireland, June 2009
Griffon vulture (Gyps fulvus) with open wings, Cebollar, Torla, Aragon, Spain, November 2008. WWE OUTDOOR EXHIBITION. NOT AVALIABLE FOR GREETING CARDS OR CALENDARS
Chaffinch (Fringilla coelebs) two males squabbling at water, Pusztaszer, Hungary, May 2008
Alpine marmot (Marmota marmota) aggressive behaviour, Hohe Tauern National Park, Austria, July 2008
Stag beetle (Lucanus cervus) two males displaying aggressive behaviour on oak tree branch at woodland edge, Suffolk, UK, Controlled conditions, July
Black grouse (Tetrao tetrix) males fighting at lek site, Cairngorms National Park, Grampian, Scotland, UK, April
Two Grey herons (Ardea cinerea) squabbling over fish fed to them by visitors, River Tame, Reddish Vale Country Park, Stockport, Greater Manchester, UK, December 2010
Two Grey herons (Ardea cinerea) on ice, squabbling over fish fed to them by visitors, River Tame, Reddish Vale Country Park, Stockport, Greater Manchester, UK, December 2010, Highly commended
Three Grey herons (Ardea cinerea) fighting in Regents Park, London, UK, April 2011
Grey seals (Halichoerus grypus) two adults fighting in the surf, Donna Nook, Lincolnshire, England, UK, October
Two common buzzards (Buteo buteo) fighting on ground in snow, Scotland, UK, January
Two Black grouse (tetrao tetrix) males fighting at lek, Deeside, Cairngorms National Park, Scotland, UK, April
Two male Alpine ibex (Capra ibex ibex) fighting in front of the Mer de Glace glacier, with a female watching, Aiguilles Rouges (Red Peaks) Regional Natural Park, Haute-Savoie, France, June
Two male Black grouse (Lyrurus tetrix) displaying at lek, Utajarvi, Finland, April
Male Panther Chameleons (Furcifer pardalis) fighting. Nosy Be, Madagascar. Photograph taken on location for BBC Wild Madagascar Series, January 2010
Red foxes (Vulpes vulpes) fighting in snowy field with woodland in the background, Vosges, France, February
Cape cobra {Naja nivea} displaying amongst vegetation, South Africa
Japanese macaque (Macaca fuscata) grinning aggressively, Jigokudani, Japan, January
Beluga whale (Delphinapterus leucas) play biting camera, Arctic circle Dive Center, White Sea, Karelia, northern Russia. Captive
African Buffalo (Syncerus caffer) males fighting, Nakuru National park, Kenya
Ruff (Philomachus pugnax) two males displaying in territorial dispute, Liminka Finland June
Two Red Foxes (Vulpes vulpes) fighting. The Netherlands. August
Great Egrets (Ardea alba) territorial fight above nest colony, Everglades National Park, Florida, USA, February
Ruffs displaying and fighting (Philomachus pugnax) Agapa River, Taimyr Peninsula, Siberia, Russia
Arctic tern (Sterna paradisaea) adult hovering directly in front of the sun, defending nest from perceived threat, Farne Islands, UK. June Non-ex
Black Rhinoceros (Diceros bicornis) charging. Etosha National Park, Namibia, January
Two male Mute swans (Cygnus olor) displaying on water in territorial dispute, UK