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Konik horse (Equus caballus) two stallions fighting, Wicken Fen, Cambridgeshire, UK, June
RF- Capercaillie (Tetrao urogallus) male displaying, Inshriach, Cairngorms National Park, Scotland, UK. February
Wild Mustang horses fighting McCullough Peaks Herd Area, Wyoming, USA
Aggressive Mute swan (Cygnus olor) Upper Bavaria, Germany, Europe
RF - Polar Bear (Ursus maritimus) males fighting, Churchill, Canada, November
Polar bear (Ursus maritimus) 3-year-olds play fighting on newly formed pack ice, Beaufort Sea, off the 1002 area of the Arctic National Wildlife Refuge, North Slope, Alaska
Booted racket tail (Ocreatus underwoodii) two males fighting in flight, Mindo, Pichincha, Ecuador
Stellers Sea Eagles (Haliaeetus pelagicus) fighting on ice floe, Hokkaido, Japan. February
White-tailed Sea Eagle (Haliaeetus albicilla) being attacked by a Common Gull (Larus canus) in flight. Norway, July. Magic Moments book plate, page 51
Two male Common cuttlefish (Sepia officinalis) compete for a female (on right) during courtship, Torbay, Devon, UK, May. 2020VISION Exhibition. 2020VISION Book Plate
Grey heron (Ardea cinerea) and Great egret (Ardea alba) fighting over fish, Lake Csaj, Pusztaszer, Hungary, December. Winner of the Portfolio category of the Terre Sauvage Nature Images Awards
Siskins (Carduelis spinus) fighting over food. Oisterwijk, The Netherlands, March
Red deer (Cervus elaphus) stags fighting at dusk during rutting season, Cheshire, UK October
Bison (Bison bison) two males head to head, Yellowstone NP, Wyoming, USA
Two Black-capped chickadees (Poecile atricapillus) in mid-air fight by a sunflower, New York, USA, February
Adult male Hippopotamuses (Hippopotamus amphibius) posturing in agressive yawn behaviour, Luangwa River, South Luangwa National Park, Zambia, November. Digital composite (two frames stitched together)
Male Greenfinches (Carduelis chloris) squabbling in flight. Dorset, UK, March
African Elephant (Loxodonta africana) bulls in dominance fight, Mono Pools national Park, Zimbabwe, Southern Africa
Head portrait of Sumatran tiger (Panthera tigris sumatrae) with face half cast in shadow, with aggressive expression, captive
Trumpeter Swans (Cygnus buccinator) in winter morning mist, showing aggression during courtship behaviour, Mississippi River, Minnesota, USA, February
Maine Coon female cat, Serafin, in fierce defensive posture
Portrait of Iberian wolf {Canis lupus sygnatus} snarling, captive, Lobo Park, Antequera, Malaga, Spain
Melanistic (black form) Leopard snarling, often called black panther (captive)
Ibex (Capra Ibex) fighting, Aiguilles Rouges National Nature Reserve, Haute-Savoie, France, June
Adult male Hippopotamus (Hippopotamus amphibius) posturing in agressive yawn behaviour, Luangwa River, South Luangwa National Park, Zambia, November
Siberian tiger {Panthera tigris altaica} two cubs play fighting, captive
Two Iberian ibex (Capra pyrenaica), adult males, fighting. Sierra Nevada National Park, Andalusia, Spain. July
Two Blue spotted mud skippers (Boleophthalmus boddarti) fighting at water's edge, Sunderban tiger reserve, West Bengal, India
Two Common terns (Sterna hirundo) in middle of aggressive encounter as one lands near nest with chick in it. Long Island, New York, USA. June
Two wild Onaqui stallions (Equus ferus) fighting for dominance in between their respective herd. Onaqui Mountain Herd Management Area, Utah, USA. September
Velvet swimming crab (Necora puber) male, in defensive posture, with wreck of the Rosalie in background, Weybourne, Norfolk, England, UK, North Sea
Bengal tiger (Panthera tigris tigris) walking through river, snarling, Bardia National Park, Terai, Nepal. Endangered
Two Red deer (Cervus elaphus) stags, fighting during the rut, Cheshire, UK. July
Two Ruffs (Philomachus pugnax) male, competing at a lek, Vardo, Norway. May
Black grouse (Tetrao tetrix) males fighting in snow during the breeding season, Viiksimo, Finland. April
Two Speckled moray eels (Gymnothorax dovii) feeding on a Sea turtle carcass on the seabed, Wolf Island, Galapagos Islands, Pacific Ocean
Two Komodo dragons (Varanus komodoensis) males, fighting on a remote beach, Komodo Island, Komodo National Park, Indonesia. Endangered
Three Spotted hyenas (Crocuta crocuta) fighting over meat from a nearby carcass with flock of White-backed vultures (Gyps africanus) watching in background, Mana Pools National Park, Zimbabwe
Two male Common / Plains zebras (Equus quagga) fighting, Okavango Delta, Botswana, Africa
Two Buzzards (Buteo buteo) fighting over pheasant carcass, Marlborough Downs, Wiltshire, UK. December
Three male Ostriches (Struthio camelus) running and flapping wings in aggressive display, Kgalagadi Transfrontier Park, South Africa
Two male Black grouse (Tetrao tetrix) fighting for territory on lek, Cairngorms National Park, Scotland, UK. May
Two male Polar bears (Ursus maritimus) standing on hind legs, sparring, Churchill, Canada. November
Two Red deer (Cervus elaphus) stags fighting, Bradgate Park, Leicestershire, UK. October
Dalmatian pelicans (Pelecanus crispus) fighting for fish. Lake Kerkini, Greece. February. Vulnerable species
Two male Black grouse (Lyrurus tetrix) fighting and sparing over territory at a snow covered lek, Vasterbotten, Sweden. April
Juvenile Southern elephant seals (Mirounga leonina) practice fighting near a King penguin (Aptenodytes patagonicus) colony, Gold Harbour, South Georgia Island
Two Grey heron (Ardea cinerea) fighting over a fish, Pusztaszer reserve, Hungary. May