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Two male Black grouse (Lyrurus tetrix) fighting and sparing over territory at a snow covered lek, Vasterbotten, Sweden. April
Juvenile Southern elephant seals (Mirounga leonina) practice fighting near a King penguin (Aptenodytes patagonicus) colony, Gold Harbour, South Georgia Island
Two Grey heron (Ardea cinerea) fighting over a fish, Pusztaszer reserve, Hungary. May
Two young male African forest elephants (Loxodonta cyclotis) fighting over small pool of water in Dzanga-Sangha National Park, Central African Republic
Alpha male Grey wolf (Canis lupus) biting angry young wolf to assert dominance, Finland. July
Male Yellowface pike-blenny (Chaenopsis limbaughi), living in abandoned tubeworm holes in sand, flaring its dorsal fin in threat display, Dominica, Eastern Caribbean
Two male Alpine Ibex (Capra ibex) fighting on a mountain slope at dawn, Varaita Valley, Alps, Italy, July
Jackdaw (Corvus monedula) pair dive bombing and harassing a Grey squirrel (Sciurus carolinensis) as it emerges from a nest box the birds want to nest in which is already occupied by the squirrel
Jackdaw (Corvus monedula) swooping down calling with claws raised to threaten a Grey squirrel (Sciurus carolinensis) in the entrance to a nest box the bird wants to nest in which is already occupied
Jackdaw (Corvus monedula) swooping in very close to threaten a Grey squirrel (Sciurus carolinensis) as it stands in the entrance to a nest box the bird wants to nest in which is already occupied by
Coots (Fulica Atra) fighting in early morning light. Valkenhorst Nature Reserve, The Netherlands, Europe. April
Dippers (Cinclus cinclus) standing on ice, ready to fight over feeding territory, Kuusamo, Finland, February
Dippers (Cinclus cinclus) fighting over feeding territory, Kuusamo, Finland, February
Male Saiga antelope (Saiga tatarica) rutting in winter, The Black Lands National Park, Kalmykia, Russia
Two male Darjeeling bush frogs (Raorchestes Annandalii) fighting over territory, Kalimpong, West Bengal, India
Moose (Alces alces) two bulls sparring in willow flat grasslands, Grand Teton National Park, Wyoming, USA. October
Greenfinch (Carduelis chloris) and Goldfinch (Carduelis carduelis) fighting in snow, Poland. February
Coots (Fulica Atra) fighting in early morning light. Valkenhorst nature reserve, Valkenswaard, The Netherlands. April
Alpine ibex (Capra ibex) adult males fighting on a steep mountain side, Valsavarenche, Gran Paradiso National Park, Aosta Valley, Italy, May
Red deer (Cervus elaphus) stags sparring in the frost. Richmond Park, London, England, UK. December
Southern elephant seals (Mirounga leonina) male fighting for breeding rights. St Andrews Bay, South Georgia Island
Golden eagle (Aquila chrysaetos), two juveniles fighting in snow. Norway. November
Coot (Fulica atra) two fighting in a territorial dispute during the breeding season on a misty morning. Valkenhorst Nature Reserve, Valkenswaard, The Netherlands, May
Two wild konik horse stallions fighting in front of herd, Oostvaardersplassen Nature Reserve, Flevoland, Netherlands
Dalmatian pelicans (Pelecanus crispus) squabbling over fish, Lake Kerkini, Greece. March
Wild-living Konik horses, (Equus caballus), fighting, Oostvaardersplassen Nature Reserve, The Netherlands, April
Mountain gorilla (Gorilla berengei), aggressive young male from Titus Group, Visoke, Volcanoes National Park, Rwanda, August
Marine iguanas (Amblyrhynchus cristatus) fighting, Punta Gavilanes, Fernandina Island, Galapagos
RF - Arctic foxes (Alopex lagopus) blue colour morphs in winter coat fighting, Hornstrandir, IcelandMarch (This image may be licensed either as rights managed or royalty free.)
Greater flamingoes (Phoenicopterus roseus) males twisting their necks together and fighting in a disused salt pan. La Tancada Salinas Nature Reserve, Ebro Delta, Catalonia, Spain
Black Grouse (Lyrurus tetrix) fighting on icem Finland. March
African elephant (Loxodonta africa) male in threat display to rival. Floor of the Ngorongoro Crater, Ngorongoro Conservation Area, Tanzania
Atlantic walruses (Odobenus rosmarus) resting on ice, with two large individuals facing off just before the one on the right stabbed the other with his tusks, Svalbard, Norway, June
Whitebacked vulture (Gyps africanus) fighting over food, Zimanga private game reserve, KwaZulu-Natal, South Africa
Southern elephant seal (Mirounga leonina), two males, equally matched with mouths open in aggression. Out of focus King penguin (Aptenodytes patagonicus) colony in background
Dalmatian wall lizard, (Podarcis melisellensis), fight between two males with distinct morphs, Croatia, April. Non-ex
Pere Davids deer (Elaphurus davidianus) two stags boxing amongst herd of hinds. Captive bred in reserve after reintroduction from Woburn Abbey, England, UK
RF - Two Red foxes (Vulpes vulpes) on hind legs play fighting, Netherlands
Canada geese (Branta canadensis) two during territorial fight in early spring, Ithaca. New York, USA
Two Buzzards (Buteo buteo) fighting, Lake Csaj, Kiskunsagi National Park, Hungary. February
Cormorants (Phalacrocorax carbo) squabbling over fish, Lake Csaj, Kiskunsagi National Park, Pusztaszer, Hungary. January
Grey herons (Ardea cinerea) fighting, Lake Csaj, Kiskunsagi National Park, Pusztaszer, Hungary. January
Grey heron (Ardea cinerea) squabbling, Lake Csaj, Kiskunsagi National Park, Pusztaszer, Hungary. February
Coastal brown bears (Ursus arctos) fighting, Lake Clarke National Park, Alaska, September
Two male Dusky groupers (Epinephelus marginatus) fighting for territory during mating season at baixa do meros, a small pinnacle at Formigas islets Protected area, near Santa Maria Island, Azores
White tailed eagles (Haliaeetus albicilla) two juveniles fighting in flight, Lake Csaj, Kiskunsagi National Park, Pusztaszer, Hungary. January
Great egret (Ardea alba) fighting over fish with Grey heron (Ardea cinerea) Lake Csaj, Kiskunsagi National Park, Pusztaszer, Hungary. January
Two wild Mustang stallions fighting in White Mountain Herd Area, Wyoming, USA. August
