Ruff (Philomachus pugnax) two males displaying in territorial dispute, Liminka Finland

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Ruff (Philomachus pugnax) two males displaying in territorial dispute, Liminka Finland

Ruff (Philomachus pugnax) two males displaying in territorial dispute, Liminka Finland June

naturepl.com / : Markus Varesvuo

Media ID 15264483

© www.naturepl.com

Aggression Brown Catalogue5 Communication Competition Contrasting Contrasts Display Face To Face Finland Males Mating Behaviour Philomachus Pugnax Rivalry Sandpipers Scandinavia Scolopacidae Territorial Two Animals Waders White

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