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Northern / Hen Harrier (Circus cyaneus) and Kestrel (Falco tinnunculus) below, fighting in flight. Vosges, France, February
African Wild Dog (Lycaon pictus) pack passing infront of large African elephant (Loxodonta africana) Mana Pools National Park, Zimbabwe October 2012
Red fox (Vulpes vulpes) cubs playing, Vosges, France, May
Mountain hares (Lepus timidus) boxing in snow, Scotland, UK, December
Wild Horses / mustangs, two stallions rearing up fighting, Pryor Mountains, Montana, USA, June
Great crested grebe (Podiceps cristatus) pair performing courtship displaying at dawn, backlit and surrounded by mist, Cheshire, UK, April
Black rhino (Diceros bicornis) and White Rhino (Ceratotherium simum) bulls facing off. Solio Game Reserve, Laikipia, Kenya. September
Two wild pinto Mustang stallions battle for dominance in Sand Wash Basin, Colorado, USA
Wild Mustang horses interacting, Pryor mountains, Montana, USA. June
Topi (Damaliscus korrigum) at sunset, Masai-Mara Game Reserve, Kenya
Male Caucasian turs (Capra caucasica) rutting with Dykh-tau Mountain in the background, Kabardino-Balkarsky Zapovednik / Reserve, Russia, July
Albino Common Iguana (Iguana iguana) captive pair from Tom Crutchfields collection, from Caribbean and South and Central America
Two Great egrets (Ardea alba) standing opposite each other with Grey heron (Ardea cinerea) in between. Lake Csaj, Pusztaszer, Hungary, January
Yellow mongoose (Cynictis penicillata), two pups nose to nose. Kalahari Gemsbok National Park, Kgalagadi Transfrontier National Park, Northern Cape, South Africa
RF -Two King penguins (Aptenodytes patagonicus) head to head, South Georgia, South Atlantic. (digitally stitched image) (This image may be licensed either as rights managed or royalty free.)
Guanaco (Lama guanicoe) female and newborn baby, nose to nose in grassland. La Pampa Province, Argentina
Southern lechwe (Kobus leche), two males about to fight, facing each other. Okavango Delta, Botswana
Head portrait of black Andalusian stallion and mare meeting for the first time in southern Spain, Europe
Royal terns (Sterna maxima) pair crossing bills during courtship behaviour, Fort DeSoto Park, Florida, USA
Zebu Cattle (Bos primigenius indicus), Los Llanos, Colombia, South America
Red Foxes (Vulpes vulpes) young play fighting. Kronotsky Zapovednik Nature Reserve, Kamchatka Peninsula, Russian Far East, July
Tibetan antelope (Pantholops hodgsoni), two males, Kekexili, Qinghai, Tibetan Plateau, China, December
Greater flamingos (Phoenicopterus ruber roseus) males displaying, lake Nakuru, Kenya
Polar bear (Ursus maritimus) 3-year-olds play fighting on newly formed pack ice, Beaufort Sea, off the 1002 area of the Arctic National Wildlife Refuge, North Slope, Alaska
RF- Little blue heron (Egretta caerulea) and Great egret (Ardea alba) in lagoon at sunrise. Fort Myers beach, Gulf Coast, Florida, USA, March
Great white egret (Ardea alba) silhouetted at sunset, Lake Csaj, Pusztaszer, Hungary, February. Winner of the Portfolio category of the Terre Sauvage Nature Images Awards competition 2015
Musk ox (Ovibos moschatus) two in habitat, Wrangel Island, Far Eastern Russia, October
Grey heron (Ardea cinerea) and Great egret (Ardea alba) fighting over fish, Lake Csaj, Pusztaszer, Hungary, December. Winner of the Portfolio category of the Terre Sauvage Nature Images Awards
Red deer (Cervus elaphus) stags fighting at dusk during rutting season, Cheshire, UK October
Two Black-capped chickadees (Poecile atricapillus) in mid-air fight by a sunflower, New York, USA, February
Adult male Hippopotamuses (Hippopotamus amphibius) posturing in agressive yawn behaviour, Luangwa River, South Luangwa National Park, Zambia, November. Digital composite (two frames stitched together)
Two Cheetahs (Acinonyx jubatus) touching noses in greeting display, Serengeti NP, Tanzania, East Africa, January
African Elephant (Loxodonta africana) bulls in dominance fight, Mono Pools national Park, Zimbabwe, Southern Africa
Two Cape Ground squirrels (Xerus inauris) greeting, Kgalagadi Transfrontier Park, South Africa Non-ex
Wild horse / mustang (Equus caballus) two young mustangs play-fighting in the Pryor Mountains, Wyoming, USA
RF - Head portrait of black Andalusian mare and stallion meeting for the first time in Spain, Europe
Gannet (Morus bassanus) breeding pair during part of their elaborate courtship ritual. Saltee Islands, Republic of Ireland, May
Unravelling Fern fronds (Unknown species). Near Nepenthes Field Camp, mid-altitude montane forest. Centre of Maliau Basin, Sabahs Lost World, Borneo
Two European goldfinches (Carduelis carduelis) fighting over food in snow, Wrocław, Lower Silesia, Poland. February
Two Great egrets (Ardea alba) face to face against white sky, Lake Csaj, Pusztaszer, Hungary. Winner of the Portfolio category of the Terre Sauvage Nature Images Awards competition 2015
Pair of Broadclub cuttlefish (Sepia latimanus) mating in a coral reef, Kagoshima Prefecture, Japan. Having succeeded in gaining the female's acceptance
New Forest donkeys, female with foal, standing on grassland, touching noses at dusk, New Forest National Park, Hampshire, England, UK. May
Two Humpback whales (Megaptera novaeangliae) nose to nose, Moorea, French Polynesia, Pacific Ocean
Pair of European bee-eaters (Merops apiaster) with courtship offering of insect prey, Pusztaszer, Kiskunsagi National Park, Hungary
Pair of Wilson's birds of paradise (Cicinnurus respublica), male performing courtship display to female from a small sapling in the middle of his display court, Batanta Island, West Papua
Two Buzzards (Buteo buteo) fighting over pheasant carcass, Marlborough Downs, Wiltshire, UK. December
Two Western grey plantain-eaters (Crinifer piscator) looking at each other, Allahein River, The Gambia
Two male Black grouse (Tetrao tetrix) fighting for territory on lek, Cairngorms National Park, Scotland, UK. May
