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A male Leafy Seadragon (Phycodurus eques) carrying eggs. Wool Bay, Edithburgh, South Australia, Gulf of St Vincent, November
Norwejian Fjord stallion running in the snow, Berthoud, Colorado, USA
Male Bullfinch (Pyrrhula pyrrhula) perched amongst Cherry blossom, Buckinghamshire, England
Head portrait of male silverback Western lowland gorilla {Gorilla gorilla gorilla} UK
Tompot blenny (Parablennius gattorugine) in bright summer mating colours, peering out from his home in a discarded tube, where he was guarding his eggs (not visible in this photo), Swanage Pier
Mountain Gorilla (Gorilla beringei beringei) Sabyinyo Group, Silverback in meadow, Volcanoes National Park, Rwanda, East Africa
Lazuli Bunting (Passerina amoena) male in breeding plumage, singing, Mono Lake Basin, California, USA. July
Asian Elephant (Elephas maximus) male, India
Asian Elephant (Elephas maximus) male, walking through forest, South India
Golden Oriole (Oriolus oriolus) male feeding its chicks at nest, Bulgaria
Hooded seal male display {Cystophora cristata} St Lawrence gulf, Canada
Wood Duck (Aix sponsa), male flapping its wings, autumn colour reflected in water, Ohio, USA
Male Redstart (Phoenicurus phoenicurus), perched on branch of flowering Hawthorn (Crataegus monogyna), Clwyd, Wales, UK, May
Mallard (Anas platyrhynchos) male flying low over water. Cambridgeshire Fens, England, March
Jaguar (Panthera onca) male walking, Pantanal, Brazil
Great crested grebes (Podiceps cristatus) performing courtship ritual at dawn, Cheshire, UK April
Male silverback Eastern lowland gorilla (Gorilla beringei graueri) in the equatorial forest of Kahuzi Biega Park, Democratic Republic of Congo, Africa, March 2009
Silverback male Eastern lowland gorilla (Gorilla beringei graueri) intimidation display by chest beating, equatorial forest of Kahuzi Biega Park, Democratic Republic of Congo, Africa, March 2009
Male Greenfinch (Carduelis chloris) amongst Pussy willow catkins, Hertfordshire, England, UK
Blackbird (Turdus merula) male, portrait, Pusztaszer, Hungary, May 2008
Red deer (Cervus elaphus) in Richmond Park with Roehampton Flats in background, London, England, UK, September. 2020VISION Book Plate
Yellowhammer (Emberiza citrinella) male perched. Scotland, UK, December
Bearded reedling / tit (Panurus biarmicus) male perched on twig, Titchwell, Norfolk, UK, June
Willow Grouse (Lagopus lagopus) male perched on snow, Finland, March. Bookplate from Danny Greens The Long Journey North
Sand lizard male {Lacerta agilis} Ainsdale NNR, Lancashire, UK
Wild Horses / mustangs, two stallions rearing up fighting, Pryor Mountains, Montana, USA, June
Booted Racket-tail hummingbird (Ocreatus underwoodii) male flying, feeding from flower, Mindo, Ecuador, Andes, South America, January
European Lynx (Lynx lynx) adult male stalking through winter birch forest, Bardu, Norway, captive
Portrait of Red deer stag (Cervus elaphus) on open moorland in snow, Cairngorms NP, Scotland, UK, December
Portrait of a grey Andalusian stallion, in Ciutadella, Menorca, Spain
Broad-tailed Hummingbird {Selasphorus platycercus} male in flight feeding on Musk Thistle (Carduus nutans) Rocky Mountain National Park, Colorado, USA, June
Highland cattle bull portrait {Bos taurus} Scotland UK
RF- Capercaillie (Tetrao urogallus) male displaying in snow, Cairngorms National Park, Scotland, February 2009
Harlequin duck (Histrionicus histrionicus) male swimming, Iceland, June 2011. Bookplate from Danny Greens The Long Journey North
Red Deer (Cervus elaphus) stag seen from behind a tree. Cairngorms National Park, Scotland, February
RF- Northern Cardinal (Cardinalis cardinalis) male perched on branch of flowering Eastern Redbud (Cercis canadensis). Dinero, Lake Corpus Christi, South Texas, USA
Acorn Woodpecker (Melanerpes formicivorus), male at granary tree showing many acorns stored for winter survival, Orange County, California, USA
Small emperor moth (Saturnia pavonia) male with wings open showing eyespots on lichen covered twig, Captive, UK, April
Satin bowerbird male at bower decorated with blue objects to attract mate, Lamington NP
Red deer stag (Cervus elaphus) calling during rut. Richmond Park, England, UK, Europe
Kirks Dik-Dik (Madoqua kirki) male standing in tall grass, Tarangire National Park, Tanzania
Male Indian peafowl / peacock (Pavo cristatus) from open forest areas on the Indian subcontinent. Captive, Jurong Bird Park, Singapore
Rear of Male African elephant {Loxodonta africana} Ngorongoro crater, Tanzania
Blue-chested hummingbird (Amazilia amabilis) male feeding from flower, Costa Rica, Central America
Indigo Bunting (Passerina cyanea) male perched, Texas, USA
TemminckAśs tragopan (Tragopan temminckii) male during courtship displayTemminck's tragopan (Tragopan temminckii) male during courtship display, showing red-blue lappets. Tibet, China
Male Giraffe-necked Weevil (Trachelophorus giraffa) Montane rainforest, Vohiparara, Ranomafana National Park, Madagascar
Male Panther Chameleon (Furcifer pardalis) stalking prey in beach side vegetation. Bay of Antongil, Masoala Peninsula National Park, north east Madagascar
Rhinoceros hornbills (Buceros rhinoceros) pair flying over the canopy with mist hanging over lowland Dipterocarp rainforest in heart of Danum Valley, Sabah, Borneo (digitally modified image)
California Quail (Callipepla californica), adult male perched on bush, acting as sentinel, Montana De Oro State Park, California, USA, July
Male Japanese macaque (Macaca fuscata) roaring to attract females during the breeding season, Jigokudani, Joshinetsu Kogen NP, Nagano, Japan
Silhouette of Red Deer (Cervus elaphus) stag calling during rut, Dyrehaven, Denmark
Sika Deer (Cervus nippon) stag in woodland, Arne RSPB Reserve, Dorset, UK, September
Sika deer (Cervus nippon) stags in summer coat, Arne RSPB Reserve, Dorset, England, August
Painted bunting (Passerina ciris) male singing, Wichita Mountains National Wildlife Refuge, Oklahoma, USA, May
Eastern / Rufous-sided towhee (Pipilo erythrophthalmus), male singing, perched amongst apple blossom in spring, New York, USA, May
Wild Turkeys (Meleagris gallopavo), two toms courting several hens, New York, USA
European river otter {Lutra lutra} male on coast, Shetland Islands, Scotland, UK
Magnificent Frigate (Fregata magnificens) male displaying by inflating his throat pouch. North Seymour, Galapagos, Ecuador, April
Male Spanish ibex (Capra pyrenaica) Sierra de Gredos, Spain, November 2008
RF- Cute tabby kitten, Stanley, aged 6 weeks, walking
Fosa / Fossa (Cryptoprocta ferox) male snarling, dry deciduous forest, Kirindy Forest, Western Madagascar, IUCN vulnerable species
African elephant (Loxodonta africana) bull with dried mud covering skin. Mud creates a monochrome texture. Savuti, Botswana. Taken on location for BBC Planet Earth series, 2005
Male Common peafowl (Pavo cristatus) displaying feathers, Ranthambore NP, Rajasthan, India
Arctic fox male portrait {Vulpes lagopus} Norway captive
RF- Adult male Serval (Felis serval) stalking prey in long grass. Serengeti National Park, Tanzania
Male Black and white colobus monkey {Colobus guereza} Lake Naivasha, Kenya, East Africa
Coppery-headed Emerald, Elvira cupreiceps, male in flight feeding on Spiral Ginger (Costus Pulverulentus), Central Valley, Costa Rica, Central America, December 2006
Male White-throated Mountain Gem (Lampornis castaneoventris). Tamalanca Mountains, Costa Rica. May
Takin male portrait {Budorcas taxicolor} Mt Qinling, Shaanxi, China
Kingfisher (Alcedo atthis) male perched in tree with mud on beak, Hertfordshire, England, UK, March
Bullocks Oriole (Icterus gabula bullockii) male in breeding plumage perched amid new leaves in spring, Mono Lake Basin, California, USA
Eastern Bluebird (Sialia sialis) male perched on flowering eastern redbud in spring, New York, USA (Digitally retouched image - flower cleanup)
Bullfinch (Pyrrhula pyrrhula) male perched on frosty twigs, Kuusamo, Finland, January
Lesser Goldfinch {Carduelis psaltria} black-backed male on Mealy sage (Salvia farinacea) Hill Country, Texas, USA
Rear view of male Bengal tiger walking {Panthera tigris tigris} Kanha NP, India - Winner in Wildlife Photographer of Year competition
Common cranes displaying by woodland {Grus grus} Finland
Male Reindeer (Rangifer tarandus) on autumn tundra. Kronotsky Zapovednik Nature Reserve, Kamchatka Peninsula, Russian Far East, October
Greenfinch (Carduelis chloris) on blackthorn / sloe berries, UK
Red deer (Cervus elaphus) stag roaring during the rut, Leicestershire, UK, October. Non-ex
Mallard drake (Anas platyrhynchos) portrait of male on water, with autumn colours reflecting. Lancashire, England, UK. November
RF- Northern Cardinal (Cardinalis cardinalis) immature male perched on Texas Red Oak (Quercus Texana). New Braunfels, San Antonio, Hill Country, Central Texas, USA
Moose (Alces alces) bull at the edge of the Great North Woods, Baxter State Park, Maine, New England, USA, Autumn
Male Greenfinch (Carduelis chloris) perching on Hawthorn, Cheshire, UK, October