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RF - Lions (Panthera leo) - two brothers patrolling territorial boundary. Short grass plains on the border of Serengeti / Ngorongoro Conservation Area (NCA) near Ndutu, Tanzania
Blacktail Prairie Dog (Cynomys ludovicianus) engaging in Jump-yip behavior - A strong arch of the back or " jump" followed by a shrill " yip" - Thought to occur when a predator
Lions (Panthera leo) - two brothers patrolling territorial boundary, affectionate behaviour, border of Serengeti / Ngorongoro Conservation Area (NCA) near Ndutu, Tanzania
Spotted-tailed quoll (Dasyurus maculatus) scent marking in Monga National Park, New South Wales, Australia. Remote camera, triggered by movement
Wild Horses / mustangs, two stallions rearing up fighting, Pryor Mountains, Montana, USA, June
Siberian tiger (Panthera tigris altaica) female territorial marking by rubbing cheek against ancient Manchurian fir tree, Land of the Leopard National Park, Far East Russia, November
Violet sabrewing hummingbird (Campylopterus hemileucurus) male hummingbirds territorial fight at dawn, Turrialba, Costa Rica
RF - Polar Bears (Ursus maritimus) males fighting, Churchill, Canada, November
Rufous-tailed hummingbird (Amazilia tzacatl) territorial fighting around Heliconia flower (Heliconia latispatha) La Selva, Costa Rica
RF- Indian eagle owl (Bubo bengalensis) head portrait, captive, from India
Wood Warbler (Phylloscopus sibilatrix) singing from perch. Wales, May
Male Caucasian turs (Capra caucasica) rutting with Dykh-tau Mountain in the background, Kabardino-Balkarsky Zapovednik / Reserve, Russia, July
Eurasian coot (Fulica atra) males fighting over territory. London, England, UK. December.;
Two Black woodpeckers (Dryocopus martius) males on tree trunk in territorial dispute, Peerdsbos, Braschaat, Belgium. April
Fosa (Cryptoprocta ferox) male marking territory onto a mating tree, Kirindy Forest, Madagascar
American avocets (Recurvirostra americana) two during aggressive (territorial) encounter, Bear River Migratory Bird Refuge, Utah, USA, May
Wood Ducks (Aix sponsa), females behave aggressively toward a male (second from right) that is trying to join them on a floating log, autumn, New York, USA, October
Hippopotamus (Hippopotamus Amphibius) territorial aggressive behavior, Chobe National Park, Botswana
Jaguar (Panthera onca) female, smelling scent marking of male. Cuiaba River, Pantanal Matogrossense National Park, Pantanal, Brazil
Red winged blackbird (Agelaius phoeniceus) male singing, Bozeman, Montana, USA. April
RF - Polar Bear (Ursus maritimus) males fighting, Churchill, Canada, November
Two male Lesser kestrels (Falco naumanni) in pursuit flight, one driving off the other from its nesting territory. Extremadura, Spain
Lioness (Panthera leo) chasing away Cheetah (Acinonyx jubatus) Serengeti / Ngorongoro Conservation Area (NCA) near Ndutu, Tanzania
European Cuckoo {Cuculus canorus} male display posture Cairngorms, Scotland. Spring
Pheasant {Phasianus colchicus} males fighting over territory, Norfolk, UK
Siskins (Carduelis spinus) fighting over food. Oisterwijk, The Netherlands, March
Alpine ibex (Capra ibex) adult male chasing two young males away, Gran Paradiso National Park, Italy, July
Red fox (Vulpes vulpes) sniffing beech tree trunk, Black Forest, Germany, Winner of Fritz Polking portfolio prize, GDT 2011 competition
Red deer (Cervus elaphus) stags fighting at dusk during rutting season, Cheshire, UK October
Bison (Bison bison) two males head to head, Yellowstone NP, Wyoming, USA
Two Black-capped chickadees (Poecile atricapillus) in mid-air fight by a sunflower, New York, USA, February
Yellow-headed Blackbird (Xanthocephalus xanthocephalus) male calling and performing courtship / territorial display in cattail marsh, Mono Lake Basin, California, USA
Male Bengal tiger (Panthera tigris tigris) known as Sundar (B2) patrolling territory, Bandhavgarh National Park, Madhya Pradesh, India. Endangered species
Asian black bear (Ursus thibetanus) scent marking tree by rubbing back on it, Land of the Leopard National Park, Russian Far East. Taken with remote camera. December
Siberian tiger (Panthera tigris altaica) sitting and scent marking tree in forest by rubbing body against it, Land of the Leopard National Park, Russian Far East. Endangered
Lion (Panthera leo) male, scent marking its territory at dawn, Ngorongoro Conservation Area / Serengeti National Park boundary, northern Tanzania
Northern parula (Parula americana) male singing from flowering crabapple (Malus sp)Northern parula (Parula americana) male singing from flowering crabapple (Malus sp.) tree in spring, near Salamanca, New York, USA, May 2022
Two male Black grouse (Lyrurus tetrix) fighting and sparing over territory at a snow covered lek, Vasterbotten, Sweden. April
Male Yellowface pike-blenny (Chaenopsis limbaughi), living in abandoned tubeworm holes in sand, flaring its dorsal fin in threat display, Dominica, Eastern Caribbean
Lions (Panthera leo) - two brothers patrolling territorial boundary at border of Serengeti / Ngorongoro Conservation Area (NCA) near Ndutu, Tanzania
Dippers (Cinclus cinclus) standing on ice, ready to fight over feeding territory, Kuusamo, Finland, February
Dippers (Cinclus cinclus) fighting over feeding territory, Kuusamo, Finland, February
Two male Darjeeling bush frogs (Raorchestes Annandalii) fighting over territory, Kalimpong, West Bengal, India
Golden damsel (Amblyglyphidodon aureus) male aggressively defends his territory on a coral reef. Piaynemo Island, Raja Ampat, West Papua, Indonesia. Tropical West Pacific Ocean
Mute swan (Cygnus olor) in a threathening territorial pose and attacking a rival Lake Geneva, near Geneva, Switzerland, March
Iberian lynx (Lynx pardinus) sniffing scent on dead branch, Parque Natural Sierra de Andujar, Andalucia, Spain. January
Male Anole Lizards (Anolis sp) in territorial display to each other. Rainforest canopy, Rio Claro Reserve, Magdalena Valley, Colombia, South America
Bengal Tiger (Panthera tigris) Arrowhead patrolling his territory, Ranthambore National Park, India
