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RF- Tawny owl with moon behind (Strix aluco), UK
Chocolate working cocker spaniel in snow, Wiltshire, UK
Grizzly bear (Ursus arctos horribilis) female with cub riding on back, Katmai National Park, Alaska, USA, August
Portrait of English Springer Spaniel (field type). Elkhorn, Wisconsin, USA, January
Working English springer spaniel puppies, age 6 weeks
Miniature American Shepherd puppy
Palomino Peruvian paso mare and foal, Sante Fe, New Mexico, USA
Cavalier King Charles Spaniels with tricolor and ruby colourations on beach, Texel, Netherlands
European hare (Lepus europaeus), Wirral, England, UK, May
Welsh Terrier, bitch with puppies aged 8 weeks in snowy scene
Suffolk Punch heavy horse in field resting head anothers back, UK, September
Black Bear (Ursus americanus) sleeping in a Sitka Spruce tree (Picea sitchensis) alongside Anan Creek, Alaska, July
RF - Australian Labradoodle, pouncing playfully
Kamchatcka brown bear (Ursus arctos beringianus) with face underwater whilst fishing for Sockeye salmon in the Ozernaya River, Kuril Lake, South Kamtchatka Sanctuary, Russia, August
RF- Quarter horses running in snow at ranch, Shell, Wyoming, USA, February
Bison (Bison bison) in snow. Yellowstone National Park, USA, February
Airedale terrier, female, running through coastal grass in autumn, Madison, Connecticut, USA. October
Pheasant (Phasianus colchicus), plumage detail. UK, March
Cavapoo puppy lying with head up
Brown-and-tan Miniature Pinscher puppy, with ears up
Proboscis monkey (Nasalis larvatus) male, Sabah, Borneo
Two Dogue de Bordeaux dogs looking over garden fence. No release available
Cavalier King Charles Spaniel, two puppies on grass, blenheim and tricolour, 5 weeks
Cavalier King Charles Spaniel, puppy with blenheim coat sitting, aged 5 months
Cavalier King Charles Spaniel, three puppies resting on cushion bed, blenheim and tricolour, 5 weeks
White-lipped snail (Cepaea hortensis) crawling over an oak sleeper retaining a garden lawn with a greenhouse in the background, Wiltshire, UK, April
RF - Brown hare (Lepus europaeus) adult in arable field, Scotland, UK, August
Two Cavapoo puppies
Red Dachshund puppy and Cavapoo puppy
Two merle Mini American Shepherd puppies
RF - Cavapoo dog, Monty, age 10 months, lying with head up
Burrowing owl (Athene cunicularia) doing rain dance, spreading wings and dancing while it rained, Arizona, USA
Burrowing owl (Athene cunicularia) rotating head, Arizona, USA
Patterdale terrier dog puppy, Korka, age 4 months
Dalmatian pelicans (Pelecanus crispus) underwater view of group feeding, Lake Kirkini, Greece. Highly commended in the GDT competition 2013
Scots pine (Pinus sylvestris) forest with Bracken (Pteridium aquilinum) ground flora in autumn, Glenfeshie, Cairngorms National Park, Scotland, UK, October
Chesapeake Bay retriever male standing in saltmarsh, Long Island Sound, Madison, Connecticut, USA
RF - Red Dachshund puppy lying down
Aerial view of Amazon Rainforest and the Yavari River, and oxbow lake, Peru. July 2015
Portrait of Skewbald Icelandic horse in the snow, Snaefellsnes Peninsula, Iceland, March
White horse of the Camargue, foal with its brown baby coat, Camargue, Southern France
Indian Cobra or Spectacled Cobra (Naja naja), Karnataka, India
Quinling Golden snub nosed Monkey (Rhinopitecus roxellana qinlingensis), adult male face portrait. Zhouzhi Nature Reserve, Qinling Mountains, Shaanxi, China
Siberian musk deer (Moschus moschiferus) male with tusks grooming, captive, Midlothian deer enclosure, UK, vulnerable species
RF- Glasswing butterfly (Greta oto), Costa Rica
RF- European brown bear (Ursus arctos) with mouth open, Kuhmo, Finland. July
Scots pine tree (Pinus sylvestris) in natural woodland, Beinn Eighe NNR, Highlands, NW Scotland, UK, May