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Meadow Brown Butterfly (Maniola jurtina) feeding from Scabious flower (Scabiosa) North Downs, Surrey, England, UK, August
RF- Tawny owl with moon behind (Strix aluco), UK
Suffolk Punch heavy horse in field resting head anothers back, UK, September
Chocolate working cocker spaniel in snow, Wiltshire, UK
Portrait of English Springer Spaniel (field type). Elkhorn, Wisconsin, USA, January
Grizzly bear (Ursus arctos horribilis) female with cub riding on back, Katmai National Park, Alaska, USA, August
Working English springer spaniel puppies, age 6 weeks
Miniature American Shepherd puppy
Cavalier King Charles Spaniels with tricolor and ruby colourations on beach, Texel, Netherlands
European hare (Lepus europaeus), Wirral, England, UK, May
Palomino Peruvian paso mare and foal, Sante Fe, New Mexico, USA
Welsh Terrier, bitch with puppies aged 8 weeks in snowy scene
Black Bear (Ursus americanus) sleeping in a Sitka Spruce tree (Picea sitchensis) alongside Anan Creek, Alaska, July
RF - Australian Labradoodle, pouncing playfully
Kamchatcka brown bear (Ursus arctos beringianus) with face underwater whilst fishing for Sockeye salmon in the Ozernaya River, Kuril Lake, South Kamtchatka Sanctuary, Russia, August
RF- Quarter horses running in snow at ranch, Shell, Wyoming, USA, February
Bison (Bison bison) in snow. Yellowstone National Park, USA, February
Walrus (Odobenus rosmarus) male resting in water, Kapp Lee, Svalbard, Norway, Arctic Ocean, July
Pheasant (Phasianus colchicus), plumage detail. UK, March
Airedale terrier, female, running through coastal grass in autumn, Madison, Connecticut, USA. October
Cavapoo puppy lying with head up
Brown-and-tan Miniature Pinscher puppy, with ears up
Proboscis monkey (Nasalis larvatus) male, Sabah, Borneo
Patterdale terrier dog puppy, Korka, age 4 months
White-lipped snail (Cepaea hortensis) crawling over an oak sleeper retaining a garden lawn with a greenhouse in the background, Wiltshire, UK, April
Verreauxs sifaka lemur (Propithecus verreauxi) dancing or skipping across open ground, Gallery Forest, Berenty Reserve, southern Madagascar
Cats whiskers / Black bat flower (Tacca chantrieri). Pollinated by flies. Cultivated in glasshouse, Surrey, England, UK. Native to Southeast Asia
Seaweeds including Thongweed / Sea thong (Himanthalia elongata), Tangleweed kelp (Laminaria digitata), Toothed wrack (Fucus serratus) and Dulse (Palmaria palmata)
RF - Brown hare (Lepus europaeus) adult in arable field, Scotland, UK, August
Two Cavapoo puppies
Red Dachshund puppy and Cavapoo puppy
Two merle Mini American Shepherd puppies
RF - Cavapoo dog, Monty, age 10 months, lying with head up
Burrowing owl (Athene cunicularia) doing rain dance, spreading wings and dancing while it rained, Arizona, USA
Burrowing owl (Athene cunicularia) rotating head, Arizona, USA
Dalmatian pelicans (Pelecanus crispus) underwater view of group feeding, Lake Kirkini, Greece. Highly commended in the GDT competition 2013
Scots pine (Pinus sylvestris) forest with Bracken (Pteridium aquilinum) ground flora in autumn, Glenfeshie, Cairngorms National Park, Scotland, UK, October
Chesapeake Bay retriever male standing in saltmarsh, Long Island Sound, Madison, Connecticut, USA
Newfoundland dog, male standing in snow. Killingworth, Connecticut, USA
RF - Red Dachshund puppy lying down
Aerial view of Amazon Rainforest and the Yavari River, and oxbow lake, Peru. July 2015
Portrait of Skewbald Icelandic horse in the snow, Snaefellsnes Peninsula, Iceland, March
An Einsiedler / Swiss warmblood mare (Equus caballus) cantering, Schwyz, Switzerland, July
White horse of the Camargue, foal with its brown baby coat, Camargue, Southern France
Bay Peruvian Paso stallion running in field, Ojai, California, USA
Cavalier King Charles Spaniel, puppy with blenheim coat sitting, aged 5 months
Two Dogue de Bordeaux dogs looking over garden fence. No release available
Cavalier King Charles Spaniel, two puppies on grass, blenheim and tricolour, 5 weeks
Chocolate Labrador retriever puppy running on garden lawn, Rhode Island, USA. May
Cockapoo puppy, aged 18 weeks, running, portrait
Serval kitten (Leptailurus serval), aged two months, near its mother, in savannah. Masai Mara national reserve, Kenya, East Africa
Long-tailed potoo (Nyctibius aethereus) camouflaged on roosting perch in lowland rainforest, Manu Biosphere Reserve, Peru
Red kangaroo (Macropus rufus) on open plain. showing brown and grey forms, Diamantina River, Queensland, Australia
Slime mould (Lamproderma scintillans) close up of sporangia growing on a Holly leaf, Buckinghamshire, England, UK, December - Focus stacked
Marbled White butterfly (Melanargia galathea) resting on flower head, Dunsdon Nature Reserve, Devon, UK. July
Inser Weiher in autumn, a pond with red coloured water, caused by dissolved tannins. Berner Seeland, Berne, Switzerland, November. Photographed for The Freshwater Project
Tibetan gazelle or Goa, (Procapra picticaudata) leaping, in mid air. Dulan Nature Reserve, Tibetan Plateau, Qinghai, China. October
Three Marbled white butterflies (Melanargia galathea) nectaring on Greater knapweed flowers (Centaurea scabiosa) in a chalk grassland meadow, Wiltshire, UK, June
Tussah moth (Antheraea godmani) close-up of head and bipectinate antennae. Santa Clara, Chiriqui Province, Panama. Focus stacked image
Eurasian red squirrel (Sciurus vulgaris) reflected in water, Cairngorms National Park, Scotland, United Kingdom, October
Group of Wild Boar (Sus Scrofa) trotting through forest of Rambouillet, near Paris, France. Autumn
Marbled white butterfly (Melanargia galathea) perched on birds foot trefoil, Volehouse Moor, Devon, UK. July
Orang utan (Pongo pygmaeus) baby sleeping in the arms of an adult, Semengoh Nature reserve, Sarawak, Borneo, Malaysia, Endangered
Adult Kea (Nestor notabilis) close-up portrait. Arthurs Pass National Park, New Zealand, July. Endangered Species
Hippopotamus (Hippopotamus Amphibius) territorial aggressive behavior, Chobe National Park, Botswana
Close up of feathers of cock Pheasant (Phasianus colchicus)
Mint moth (Pyrausta aurata) feeding on Wild marjoram flower (Origanum vulgare), chalk grassland meadow, Wiltshire, UK, July
Chocolate labrador retriever, two puppiessitting in spring flowers. Haddam, Connecticut, USA. June
RF - Cavapoo dog, Monty, 10 months, in play-bow stance
Goldendoodle puppy with raised paw
Giant peacock moth (Saturnia pyri) Sierra de Grazalema Natural Park, Ubrique, southern Spain, April
Horse (Equus caballus) grazing on thistles, UK
Hoar frost on Beech leaves (Fagus sylvatica) Rydal, Lake District, England, UK. December 2010
RF- Icelandic horse showing flehmen response, Iceland. July
Black Merens stallion running in pasture. Northern France, Europe. February
Bay Icelandic horse feeding in the snow, Snaefellsnes Peninsula, Iceland, March
Bay Icelandic horse trotting in the snow, Snaefellsnes Peninsula, Iceland, March
RF- Pinto Quarter horse in snow at ranch, Shell, Wyoming, USA, February
Comtois Horse (Equus caballus). Camargue reserve, Rhone, France, September
Flightless cormorant (Phalacrocorax harrisi) pair in courtship Puerto Pajas, Isabela Island, Galapagos
Golden eagle (Aquila chrysaetos) male head portrait, Kalvtrask, Vasterbotten, Sweden. December
Cute chocolate Toy Goldendoodle puppy in play-bow
Cute red and red-and-white Cavapoo puppies, 5 weeks, hugging, against white background