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Chocolate working cocker spaniel in snow, Wiltshire, UK
Suffolk Punch heavy horse in field resting head anothers back, UK, September
RF- Tawny owl with moon behind (Strix aluco), UK
Grizzly bear (Ursus arctos horribilis) female with cub riding on back, Katmai National Park, Alaska, USA, August
Portrait of English Springer Spaniel (field type). Elkhorn, Wisconsin, USA, January
Working English springer spaniel puppies, age 6 weeks
Meadow Brown Butterfly (Maniola jurtina) feeding from Scabious flower (Scabiosa) North Downs, Surrey, England, UK, August
Miniature American Shepherd puppy
RF - Australian Labradoodle, pouncing playfully
Cavalier King Charles Spaniels with tricolor and ruby colourations on beach, Texel, Netherlands
Kamchatcka brown bear (Ursus arctos beringianus) with face underwater whilst fishing for Sockeye salmon in the Ozernaya River, Kuril Lake, South Kamtchatka Sanctuary, Russia, August
RF- Quarter horses running in snow at ranch, Shell, Wyoming, USA, February
Bison (Bison bison) in snow. Yellowstone National Park, USA, February
Palomino Peruvian paso mare and foal, Sante Fe, New Mexico, USA
Black Bear (Ursus americanus) sleeping in a Sitka Spruce tree (Picea sitchensis) alongside Anan Creek, Alaska, July
Pheasant (Phasianus colchicus), plumage detail. UK, March
European hare (Lepus europaeus), Wirral, England, UK, May
Welsh Terrier, bitch with puppies aged 8 weeks in snowy scene
Airedale terrier, female, running through coastal grass in autumn, Madison, Connecticut, USA. October
Walrus (Odobenus rosmarus) male resting in water, Kapp Lee, Svalbard, Norway, Arctic Ocean, July
Brown-and-tan Miniature Pinscher puppy, with ears up
Patterdale terrier dog puppy, Korka, age 4 months
Chocolate Labrador retriever puppy running on garden lawn, Rhode Island, USA. May
Long-tailed potoo (Nyctibius aethereus) camouflaged on roosting perch in lowland rainforest, Manu Biosphere Reserve, Peru
White-lipped snail (Cepaea hortensis) crawling over an oak sleeper retaining a garden lawn with a greenhouse in the background, Wiltshire, UK, April
Verreauxs sifaka lemur (Propithecus verreauxi) dancing or skipping across open ground, Gallery Forest, Berenty Reserve, southern Madagascar
Tussah moth (Antheraea godmani) close-up of head and bipectinate antennae. Santa Clara, Chiriqui Province, Panama. Focus stacked image
Cats whiskers / Black bat flower (Tacca chantrieri). Pollinated by flies. Cultivated in glasshouse, Surrey, England, UK. Native to Southeast Asia
Eurasian red squirrel (Sciurus vulgaris) reflected in water, Cairngorms National Park, Scotland, United Kingdom, October
Group of Wild Boar (Sus Scrofa) trotting through forest of Rambouillet, near Paris, France. Autumn
Seaweeds including Thongweed / Sea thong (Himanthalia elongata), Tangleweed kelp (Laminaria digitata), Toothed wrack (Fucus serratus) and Dulse (Palmaria palmata)
RF - Brown hare (Lepus europaeus) adult in arable field, Scotland, UK, August
Two Cavapoo puppies
Red Dachshund puppy and Cavapoo puppy
Two merle Mini American Shepherd puppies
RF - Cavapoo dog, Monty, age 10 months, lying with head up
Burrowing owl (Athene cunicularia) doing rain dance, spreading wings and dancing while it rained, Arizona, USA
Burrowing owl (Athene cunicularia) rotating head, Arizona, USA
Dalmatian pelicans (Pelecanus crispus) underwater view of group feeding, Lake Kirkini, Greece. Highly commended in the GDT competition 2013
Mint moth (Pyrausta aurata) feeding on Wild marjoram flower (Origanum vulgare), chalk grassland meadow, Wiltshire, UK, July
Cavapoo puppy lying with head up
Proboscis monkey (Nasalis larvatus) male, Sabah, Borneo
Cavalier King Charles Spaniel, puppy with blenheim coat sitting, aged 5 months
Two Dogue de Bordeaux dogs looking over garden fence. No release available
Cavalier King Charles Spaniel, two puppies on grass, blenheim and tricolour, 5 weeks
RF - Five Cavapoochon puppies, age 6 weeks, sitting in a line
Cockapoo puppy, aged 18 weeks, running, portrait
Cockapoo puppy, aged 4 months, running, portrait
Red deer stag (Cervus elaphus) with female during rut, Exmoor National Park, Somerset / Devon, England. November
Serval kitten (Leptailurus serval), aged two months, near its mother, in savannah. Masai Mara national reserve, Kenya, East Africa
Female Serval (Leptailurus serval) portrait. Masai Mara national reserve, Kenya, East Africa
Marbled white butterfly (Melanargia galathea) sequence showing landing and taking off from a Greater knapweed (Centaurea scabiosa) flower. Yorkshire, UK. Digital composite
Red kangaroo (Macropus rufus) on open plain. showing brown and grey forms, Diamantina River, Queensland, Australia
King penguin (Aptenodytes patagonicus) chicks gathered in a creche, Fortuna Bay, South Georgia Island
Slime mould (Lamproderma scintillans) close up of sporangia growing on a Holly leaf, Buckinghamshire, England, UK, December - Focus stacked
Slime mould (Trichia botrytis) sporangia starting to split to release spores, on underside of rotting Oak (Quercus sp) log. Hertfordshire, England, UK. December. Focus stacked image
RF - Red deer (Cervus elaphus) stag standing amongst Scots pine (Pinus sylvestris) tree trunks in forest. Cairngorms National Park, Scotland, UK. January
RF - Marbled white butterfly (Melanargia galathea), roosting on dew covered Bent grass (Agrostis sp) at dawn. Devon, England, UK. July
Marbled white butterfly (Melanargia galathea) roosting among tall grasses at sunrise and bathed in warm, early morning sunlight, Volehouse Moor, Devon, UK. July
Marbled White butterfly (Melanargia galathea) resting on flower head, Dunsdon Nature Reserve, Devon, UK. July
Marbled white butterfly (Melanargia galathea) on Wild marjoram (Origanum vulgare) Farthing Downs, Surrey, England, July
Inser Weiher in autumn, a pond with red coloured water, caused by dissolved tannins. Berner Seeland, Berne, Switzerland, November. Photographed for The Freshwater Project
Common bee fly (Bombylius major) nectaring on a Greater periwinkle flower (Vinca major), Wiltshire garden, UK, April
Tibetan gazelle or Goa, (Procapra picticaudata) leaping, in mid air. Dulan Nature Reserve, Tibetan Plateau, Qinghai, China. October
Three Marbled white butterflies (Melanargia galathea) nectaring on Greater knapweed flowers (Centaurea scabiosa) in a chalk grassland meadow, Wiltshire, UK, June
Elephant yam flower (Amorphophallus paeoniifolius) spadix surrounded by spathe. Cultivated in glasshouse
Comma butterfly (Polygonia c-album) camouflaged amongst dead vegetation. Peak District National Park, Derbyshire, UK, September
Marbled white butterfly (Melanargia galathea) perched on birds foot trefoil, Volehouse Moor, Devon, UK. July
Orang utan (Pongo pygmaeus) baby sleeping in the arms of an adult, Semengoh Nature reserve, Sarawak, Borneo, Malaysia, Endangered
Adult Kea (Nestor notabilis) close-up portrait. Arthurs Pass National Park, New Zealand, July. Endangered Species
Blue morpho (Morpho helenor) butterflies feeding on mango fruit, butterfly observatory, La Paz Waterfall Gardens, Costa Rica
Hippopotamus (Hippopotamus Amphibius) territorial aggressive behavior, Chobe National Park, Botswana
Marbled white butterfly (Melanargia galathea) on knapweed flower, Aosta Valley, Gran Paradiso National Park, Italy
Bairds tapir (Tapirus bairdii) resting in mud wallow in Corcovado National Park, Costa Rica, May. Endangered
Meadow brown (Maniola jurtina) resting on grass in Cohens Field, Hampstead Heath, London, England, UK, June
Close up of feathers of cock Pheasant (Phasianus colchicus)
Meadow Brown Butterfly (Maniola jurtina) female basking, wings open, Southwest London, UK, July
Wall brown butterfly (Lasiommata megera), Lorsch, Hessen, Germany. June. Meetyourneighbours.net project
Marbled white butterfly (Melanargia galathea) resting on grass head, backlit against black background, Dunsdon, Devon, UK. July 2014
Cute chocolate Toy Goldendoodle puppy in play-bow
Cute red and red-and-white Cavapoo puppies, 5 weeks, hugging, against white background
Cavalier King Charles Spaniel, three puppies resting on cushion bed, blenheim and tricolour, 5 weeks
Cheetah cub (Acinonyx jubatus) climbing up tree stump. Masai Mara national reserve, Kenya, East Africa