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Great grey owl (Strix nebulosa) preparing to land, in tree, Yellowstone National Park, Wyoming, USA
A female kingfisher (Alcedo atthis) fishing, flying out of an ice hole in winter. Leeds, Yorkshire, UK. January
Emerald damselfy (Lestes sponsa) at Ledstone pond near Leeds, UK. September. Focus stacked
Firecrest (Regulis ignicapilla) in flight, Parainen, Uto, Finland. October
Greenfinch (Carduelis chloris) and Goldfinch (Carduelis carduelis) fighting in snow, Poland. February
Black-headed gull (chroicocephalus ridibundus) diving into water, hunting fish / shellfish at sunrise. Baltic Sea, Poland. Sequence 1 of 2
White stork (Ciconia ciconia) flying back to its nest in Oak (Quercus robur) tree to feed its three large chicks, Knepp Estate, Sussex, UK, June
Looking down on Fulmar (Fulmarus glacialis) soaring above choppy sea, Orkney, Scotland, UK, May
Fulmar (Fulmarus glacialis) in flight against sky, Orkney, Scotland, UK, May
Kestrel (Falco tinnunculus), juvenile, with its prey on fence post in late evening light. Hampshire, UK, June
Common buzzard (Buteo buteo) perched on stump with wings spread, Northamptonshire, UK, January
RF - Red admiral butterfly (Vanessa atalanta) feeding on rotten apples. Cornwall, England, UK. October. (This image may be licensed either as rights managed or royalty free.)
Shelduck (Tadorna tadorna) landing on water. Gloucestershire, England, UK. March
Lantern bug (Pyrops whiteheadi), detail of wing. Danum Valley, Sabah, Borneo
Campbell albatross (Thalassarche impavida), flying off Campbell Island (seen in background), January
Marsh fritillary butterfly (Euphydryas aurinia) nectaring on a Meadow buttercup (Ranunculus acris) flower in a chalk grassland meadow, Wiltshire, UK, June
Close up of the spots pattern on wing of a Giant owl butterfly (Caligo sp) Mindo, Ecuador
Buzzard (Buteo buteo) landing in rain, Lorraine, France, March
Northern harrier (Circus hudsonius), two juvenile females hunting. North Park, Colorado, USA. August
Mallard (Anas platyrhynchos) shaking its wings, Martin Mere WWT Reserve, Lancashire, England, UK
Saddle-billed stork (Ephippiorhynchus senegalensis) flapping wings after preening. Liuwa Plain National Park, Zambia
Black-eared kite, (Milvus migrans lineatus), Bi Tan Bay, Taiwan
Great Grey Owl (Strix nebulosa) flying directly overhead, Kuhmo Finland February
Golden Eagle adult (Aquila chrysaetos) silhouetted in tree after sunset, Kuusamo, Finland, January
Mallard (Anas platyrhynchos) shaking its wings with slow shutter speed to reveal movement, Martin Mere WWT Reserve, Lancashire, UK
Red-footed booby (Sula sula) landing among salt bushes, looking for nesting materials, with Frigate birds behind, Genovesa Island, Galapagos
Nazca booby (Sula granti) in flight at dusk. Punta Suarez, Espanola Island, Galapagos
Blue-footed booby (Sula nebouxii) on sea, tossing fish. Turtle Cove, Santa Cruz Island, Galapagos
Canada goose (Branta canadensis) with gosling, a few days old. Massachusetts, USA. April
Atlantic puffin (Fratercula arctica) in breeding plumage flapping wings on sea cliff top in seabird colony, Scotland, UK, May. Digital composite
Hawk Owl (Surnia ulula) hunting, Kuhmo, Finland, March
Common redstart (Phoenicurus phoenicurus) taking off, Sierra de Grazalema Natural Park. Southern Spain. September
RF - Andean condor (Vultur gryphus) flying over Torres del Paine Massif. Torres del Paine National Park, Chilean Patagonia, Chile. March
White capped albatross (Thalassarche steadi) in flight at sea. Auckland Islands (Subantarctic), New Zealand, February
Red-legged bumble-moth (Homoeocera gigantea) on Lichen covered tree trunk. Costa Rica
Tricoloured heron (Egretta tricolor) fishing by flying low over water, amongst Water lilies. Everglades National Park, Florida, USA. March
Galapagos hawks (Buteo galapagoensis) in flight, Galapagos, Ecuador. Vulnerable species
Intermediate egret (Egrettta intermedia), Keoladeo NP, Bharatpur, India
Female Meadow brown butterfly (Maniola jurtina) feeding on thistle, Lewisham, London, UK, August
Blue tit (Parus caeruleus) perched on tree trunk, looking at camera with wing stretched. Haukipudas, Finland. January
Striated caracara (Phalcoboenus australis) group with one landing from above with only wings and tail visible. Carcass Island, Falkland Islands, October
Egytian Goose (Alopochen aegyptiaca) rear view bathing, sings spread, Zimanga Private Game Reserves, South Africa
Sandhill crane (Grus canadensis) landing at sunset. North America. Highly honoured in the Art in Nature category of the Natures Best Photography Windland Smith Rice International Awards Competition
RF - Spicebush swallowtail butterfly (Papilio troilus) nectaring on Zinnia in farm garden, wild and free. Madison, Connecticut, USA
RF - Camouflaged moth (Agathia codina) Kinabalu National Park, Sabah, Malaysian Borneo. September
RF - Monarch butterfly (Danaus plexippus) with life-cycle tied to the presence of milkweed plants (Asclepias sp). East Haddam, Connecticut, USA. September
Saturniid moth (Saturniidae) with wings open to reveal eyespots, a means of deterring predators, Caribbean Slopes, Cordilera de Talamanca, Costa Rica
Male Common copper (Lycaena salustius) resting with wings open, Cape Kidnappers, Hawkes Bay, New Zealand, November
