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RF - Marabou (Leptoptilos crumenifer) with Whitebacked vultures (Gyps africanus) Zimanga Private Game Reserve, KwaZulu-Natal, South Africa
Straw-coloured fruit bat (Eidolon helvum), female flying carrying pup on front. Lamin, Gambia
Chinese Sparrowhawk (Accipiter soloensis) flying Guangshui, Hubei province, China. July
Peacock mantis (Pseudempusa pinnapavonis) in defensive posture; captive occurs in Burma and Thailand
Great pied hornbill (Buceros bicornis) bird photographed in flight in Hong Bung He, Dehong, Yunnan, China
Pheasant (Phasianus colchicus) male calling, East Frisian Islands, Wittbulten National Park. Wadden Sea UNESCO World Heritage Site, Germany, June
RF- Glasswing butterfly (Greta oto), Costa Rica
RF- Banded demoiselle (Calopteryx splendens), resting on dew covered reed, Lower Tamar Lakes, Cornwall/Devon border, UK. May
RF- Bannded demoiselle (Calopteryx splendens), resting on buttercup, Lower Tamar Lakes, Cornwall/Devon border, UK. May
RF- Chaffinch (Fringilla coelebs) male in flight in snow. Glenfeshie, Scotland, February
Peregrine falcon (Falco peregrinus), adult male in flight. Bristol, UK. March
Lilac-breasted roller (Coracias caudata) stretching wings on acacia branch, Masai Mara Game Reserve, Kenya. November
Golden Eagle (Aquila chrysaetos) in flight rear view, Kuusamo, Finland, February
Male Ostrich (Struthio camelus) protecting chicks from the sun with its wings, Samburu National Reserve, Kenya, Africa
Great Grey Owl (Strix nebulosa) landing in snow, Kuusamo, Finland
Long eared owl (Asio otus) in flight, taking off from oak tree snag. Niedersechsische Elbtalaue Biosphere Reserve, Elbe Valley, Lower Saxony, Germany
Snowy owl (Bubo scandiacus) in flight, Boundary Bay, British Columbia, Canada. February. First place in the Overall Category of the 2013 Great Backyard Bird Count photo competition
Steppe eagle (Aquila nipalensis) in flight, Oman, November
RF- White tailed sea eagle (Haliaeetus albicilla) in flight, Flatanger, Norway, June
RF- Golden eagle (Aquila chrysaetos) landing in snow, Flatanger, Norway. November
Small copper butterfly (Lycaena phlaeas) in meadow. Peak District National Park, Derbyshire, UK. May
RF - Female emperor dragonfly (Anax imperator) laying eggs on garden pond, Broxwater, Cornwall, UK. July
Emperor moth (Saturnia pavonia) female wide angle view showing heather moorland habitat. Peak District National Park, UK. April
Common tern (Sterna hirundo) in flight over water, Hampstead Heath, London, UK, August
Puffin (Fratercula arctica) standing on Sea thrift (Armeria maritima) with wings out stretched with a beak full of sand eels. Close-up portrait. Fair Island, Shetland Islands, Scotland, July
Orange Tip Butterfly (Anthocharis cardamines) male resting on Bluebell (Endymion / Hyacinthoides nonscriptus). Peak DIstrict National Park, Derbyshire, UK. April
Glorious scarab (Chrysina gloriosa) flying, Brewster County, Texas, USA. Controlled conditions. August
Barn Owl (Tyto alba) hunting / hovering, Somerset, UK, trained birdBarn Owl (Tyto alba) hunting/hovering, Somerset, UK, trained bird
Golden eagle (Aquila chrysaetus) landing, Kuusamo, Finland, January
Whooper Swan (Cygnus cygnus) with its beak in its feathers. The Netherlands, January
Caribbean flamingo (Phoenicopterus ruber) flock flying in formation, Ria Celestun Biosphere Reserve, Yucatan Peninsula, Mexico, January. Bookplate
Banded demoiselle (Calopteryx splendens) group of four on plant stems, River Leijgraaf, Nijmegen, the Netherlands, August 2013
Great grey owl (Strix nebulosa) taking off from snow, Kuhmo, Finland, February
Japanese / Red-crowned crane (Grus japonicus) three in flight, Hokkaido Japan February
Swallowtail butterfly (Papilio machaon), wings open, resting on plant stem. The Netherlands. August
A female kingfisher (Alcedo atthis) fishing through an ice hole in winter. Leeds, Yorkshire, UK. January. Sequence
Male Black grouse (Tetrao / Lyrurus tetrix) displaying at lek. Kuusamo, Finland, April
RF - Ruppell's griffon vulture (Gyps rueppellii) drying its wings after a rain storm, perched on strangely gnarled tree (known locally as the Devil Tree')
Black Grouse (Tetrao tetrix) males at lek, Utajarvi, Finland, April
Kingfisher (Alcedo atthis) diving into water to catch fish, Near Bourne, Lincolnshire, England, UK. January
Bald eagle (Haliaeetus leucocephalus) flying over water in falling snow. Kachemak Bay, Alaska, USA. February
Whooper swans (Cygnus cygnus) pair engaged in courtship as snow falls, Myvatn lake, Iceland, March
Namib desert dune ant (Camponotus detritus), queen on sand with visible wing scars on her thorax, Swakopmund, Namibia
Glasswing butterfly (Greta oto) resting on leaf. Captive
Ghost bat (Macroderma gigas) in flight, Pine Creek, Northern Territory, Australia
Sparrowhawk (Accipiter nisus) juvenile flying, Parainen Uto, Finland. September
Goshawk (Accipiter gentilis) flying in snow, Mainua Kajaani, Finland. March
European bee-eater (Merops apiaster) two perched on branch, Lower Silesia, Poland
