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Peacock katydid (Pterochroza ocellata) on leaf in defensive posture, Yasuni National Park, Amazon Basin, Ecuador
Two Canada geese (Branta canadensis) in flight with autumn foliage behind, Maryland, USA. October
Red grouse (Lagopus lagopus) coming in to land on flowering Heather (Calluna vulgaris), Yorkshire, UK. September
Male Banded demoiselle (Calopteryx splendens), abstract study of wings and body, Tamar Lakes, Cornwall, UK. June
Swallowtail butterfly (Papilio machaon) in flight, Finland, June
Welcome swallow (Hirundo neoxena) in flight over pond. Christchurch, New Zealand
Juvenile Kea (Nestor notabilis) in flight. Arthur's Pass National Park, South Island, New Zealand. Endangered Species
Common tern (Sterna hirundo) adult in breeding plumage flying, carrying fish for young, Long Island, New York, USA, June
Redstart (Phoenicurus phoenicurus) juvenile male, in flight, chasing an insect, Parainen Uto, Finland. September
Nutcracker (Nucifraga caryocatactes) in flight with wings spread, Sipoo, Finland. January
Wood sandpiper (Tringa glareola) in flight, Parainen Uto, Finland. August
Hen harrier (Circus cyaneus) juvenile male, in flight, Parainen Uto, Finland. September
Two Buzzards (Buteo buteo) fighting over pheasant carcass, Marlborough Downs, Wiltshire, UK. December
Male Eastern bluebird (Sialia sialis) feeding on snow-covered Holly (Ilex sp) berries, near Ithaca, New York, USA. January
Male Giant kingfisher (Megaceryle maxima) diving, Allahein River, The Gambia
Common greenshank (Tringa nebularia) wading in river with wings spread, Allahein River, The Gambia
Spoonwinged lacewing (Nemoptera bipennis) resting on a flower, Sierra de Guadarrama National Park, Madrid, Spain. June
Grey-headed flying-fox (Pteropus poliocephalus) in flight, with tongue out after licking water from its body. Having just completed a high-speed belly dip into a river on a hot day to wet its belly
Two male Black grouse (Tetrao tetrix) fighting for territory on lek, Cairngorms National Park, Scotland, UK. May
Soft focus close up of male Banded demoiselle damselfly (Calopteryx splendens) body and wings, roosting among grasses. Cornwall, UK. July
Dalmatian pelicans (Pelecanus crispus) fighting for fish. Lake Kerkini, Greece. February. Vulnerable species
Male Black grouse (Tetrao tetrix) performing courtship display at lek, Cairngorms National Park, Scotland, UK. April
Great egret (Ardea alba) standing on a branch looking under its wing, Big Cypress National Preserve, Florida, USA
Great blue heron (Ardea herodias) taking flight, Acadia National Park, Maine, USA. June
Little owl (Athene noctua) taking flight, NorthYorkshire, UK. June, 2021
Two Grey heron (Ardea cinerea) fighting over a fish, Pusztaszer reserve, Hungary. May
Eurasian hoopoe (Upupa epops) in flight with a grub in its beak, Pusztaszer reserve, Hungary. May
Black stork (Ciconia nigra) fishing, Pusztaszer reserve, Hungary. May
Pair of Eurasian bee-eaters (Merops apiaster) perched on branch, Pusztaszer reserve, Hungary. May
European roller (Coracias garrulus) in flight, Pusztaszer reserve, Hungary. May
European turtle dove (Streptopelia turtur) in flight, Pusztaszer reserve, Hungary. May
Four spotted chaser (Libellula quadrimaculata) on Common heather (Calluna vulgaris), Klein Schietveld Nature Reserve, Brasschaat, Belgium. August
Western reef egret (Egretta gularis) running through shallow water with wings open, chasing small fish, Kotu creek, The Gambia
Willowherb hawkmoth (Proserpinus proserpina) resting on a leaf, Gavarnie, South of France. June
Pamir bee hawkmoth (Hemaris ducalis) resting on a leaf, Chatkal Mountainss, Dzhalal-Abad region, Kyrgyzstan
Green-veined white butterfly (Pieris napi) resting on a flower, Tandragee, County Armagh, Northern Ireland. May
Male Spanish moon moth (Graellsia isabellae) close up of forewing eyespot, Switzerland
Two Clouded magpie moth (Abraxas sylvata) resting on a fern frond, Gosford Forest Park, County Armagh, Northern Ireland. June
Queen butterfly (Danaus gilippus) expanding wings after emerging from chrysalis on Aquatic milkweed (Asclepias perennis). Hill Country, Texas, USA. Sequence 12/12
Buff-necked ibis (Theristicus caudatus) with wings outstretched, Pocone, Brazil
Ventral view of Swallowtail butterfly (Papilio machaon) in flight. The Netherlands. July
Hills sheathtail bat (Taphozous hilli) flying out of an abandoned mine in late evening, Tennant Creek, Northern Territory, Australia
Eastern horseshoe bat (Rhinolophus megaphyllus) flying out from an abandoned mine in late evening. Its orange colouring is due to bleaching from the intense ammonia atmosphere in the mine
Robin (Erithacus rubecula) carrying a red berry in its beak as it takes off, Parainen Uto, Finland. October
Galapagos flightless cormorant (Phalacrocorax harrisi) drying stunted wings after fishing, Fernandina Island, Galapagos, South America
Mute swan (Cygnus olor) landing on frozen lake, Richmond Park, London, UK. February
Common tern (Sterna hirundo) catching a mayfly, Hjalstaviken, Uppland, Sweden. Nordic Nature Photo Competition 2021 Highly Commended - Nordic Bird Photo of the Year category
Great white egret (Ardea alba) diving to catch prey in shallow pond, Valkenhorst Nature Reserve, The Netherlands, August
