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Barn owl (Tyto alba) in flight, hunting, Hampshire, England, UK. Captive
Golden Eagle (Aquila chrysaetos) juvenile in flight, Norway, November
Jay (Garrulus glandarius), two fighting in mid-air with another observing. Norway. October
Wandering albatross (Diomedea exulans) in flight over the ocean, Drake Passage, Southern Ocean
Atlantic puffin (Fratercula arctica) landing, wings outstretched. Machias Seal Island, off the coast of Maine, USA. July
White-tailed eagle (Haliaeetus albicilla) with backlit wings, and water spray, Norway, October
Female Harpy Eagle (Harpia harpyja) in flight. Returning to its nest. Pousada Currupira d Araras, south west Brazil
Buzzard (Buteo buteo) in flight, portrait, Marlborough Downs, Wiltshire, UK. December
Calleta silkmoth (Eupackardia calleta), Texas USA Controlled conditions
Marsh fritillary butterfly (Euphydryas aurinia) nectaring on a Meadow buttercup (Ranunculus acris) flower in a chalk grassland meadow, Wiltshire, UK, June
Japanese crane (Grus japonensis) displaying, wings in heart shape, Hokkiado, Japan, February
Blue-footed booby (Sula nebouxii) landing, South coast, Santa Cruz Island, Galapagos
Red-billed tropicbird (Phaethon aethereus) in flight, Plazas Island, Galapagos
RF - Apollo butterfly (Parnassius apollo) nectaring on flower. North Tyrol, Austria. June. Focus stacked image. (This image may be licensed either as rights managed or royalty free.)
Red-footed booby (Sula sula) in flight with wings outstretched. Genovesa Island, Galapagos
Small pearl-bordered fritillary butterfly (Boloria selene), Marsland mouth, Cornwall, UK. JUne 2016
Atlantic Puffins (Fratercula arctica) flying near cliff top, Isle of Lunga, Isle of Mull, Treshnish Isles, Scotland, UK, June
Robin (Erithacus rubecula) feeding on Gelder Rose berries, Germany. January, January
Whooper swans (Cygnus cygnus) two in flight, during snowfall, Lake Kussharo, Japan, February
White tailed eagle (Haliaeetus albicilla) calling in flight, Isle of Mull, Argyll Scotland, UK. October
Common loon (Gavia immer), in breeding plumage, running across water to take flight, Michigan, USA. June
Rhinoceros hornbill (Buceros rhinoceros) male inflight, carrying mouse prey back to nest, Budo-Sungai Padi National Park, Thailand
Ghost bat (Macroderma gigas) in flight, Pine Creek, Northern Territory, Australia
Great grey owl (Strix nebulosa) hunting over snow, woodland in background. Kuusamo, Finland. February
Osprey (Pandion haliaetus) carrying fish above river in Taipei, Taiwan
Red Bird-of-Paradise (Paradisaea rubra) male performing practice display at tree-top lek, Raja Ampat, Indonesia. *Medium repro only
Golden eagle (Aquila chrysaetos) Sodankyla, Lokka, Finland, April
Saturniid moth (Leucanella hosmera), Chiriqui Province, Panama, South America
Sparrowhawk (Accipiter nisus) adult female, Norway, October
Grey-headed flying-fox (Pteropus poliocephalus) spotting and flaring wings to come in to land on a branch. Yarra Bend Park, Kew, Victoria, Australia. ÔÇ¿November 2019
Cuckoo (Cuculus canorus) in flight, Germany, April. May
Little red flying fox (Pteropus scapulatus) female and suckling baby in flight, searching for place to roost. Nitmiluk National Park, Northern Territory, Australia
Grey-headed flying-fox (Pteropus poliocephalus) hanging from a branch with one wing spread, Yarra Bend Park, Kew, Victoria, Australia.ÔÇ¿
Blue-footed booby (Sula nebouxii) diving towards sea, folding wings. Northeast coast, Santiago Island, Galapagos. Sequence 1/2
Blue-footed booby (Sula nebouxii) landing, Santa Cruz Island, Galapagos, Ecuador
Common buzzard (Buteo buteo) landing on prey, wings open, Scotland, UK, September 2016
Sarus crane (Grus antigone), male displaying, Keoladeo NP, Bharatpur, India
Group of Tissue moths (Triphosa dubitata) hibernating in a limestone cave. Peak District National Park, Derbyshire, UK. November
Great white egret (Egretta Alba) preening its feathers, Valkenhorst Nature Reserve, Valkenswaard, The Netherlands, July
Golden eagle (Aquila chrysaetos) in flight over snow. Kalvtrask, Vasterbotten, Lapland, Sweden. January
Eastern bluebird (Sialia sialis) male fluttering wings while perched on cherry (Prunus sp.) branch in spring, New York, USA, April
Snowy Egret (Egretta thula) adult in breeding plumage taking flight, Viera Wetlands, Florida, USA, March
Horned puffins (Fratercula corniculata), one landing with outspread wings, to join its mate on cliff ledge, St. Paul Island, Pribilofs, Alaska, USA, July
Indian Runner Duck, flapping wings
Osprey (Pandion haliaetus) fishing. Rothiemurchus, Cairngorms National Park, Scotland, UK
Rivolis hummingbird (Eugenes fulgens) nectaring on Fuchsia (Fuchsia sp) flower. Costa Rica
Burrowing owl (Athene cunicularia) doing rain dance, spreading wings and dancing while it rained, Arizona, USA
Lammergeier / Bearded vulture (Gypaetus barbatus) flying, Leukerbad, Valais, Switzerland, February. Highly commended in the Portfolio category of the Terre Sauvage Nature Images Awards 2017
Great white egret (Ardea alba) group of three in winter, Kiskunsag National Park, Hungary. Winner of the Bird Category of Nature Photographer of the Year Awards 2018
Chinese moon moth (Actias dubernardi, female at rest amongst leaves. Dayaoshan, Jinxin, Guangxi, China
Shag (Phalacrocorax aristotelis) in flight over sea. Great Saltee Island, Saltee Islands, County Wexford, Republic of Ireland. June
Oleander Hawkmoth (Daphnis nerii) resting, Southern Sicily, Italy. March
Snowy owl (Bubo scandiacus) in flight, Ontario, Canada, January
Wild Buffons macaw x Scarlet macaw (Ara ambiguus x Ara macao) hybrid, Costa Rica
Fulvous forest skimmer, or Russet percher dragonfly (Neurothemis fulvia) female, Sai Kung, Hong Kong, China
RF - Marabou (Leptoptilos crumenifer) with Whitebacked vultures (Gyps africanus) Zimanga Private Game Reserve, KwaZulu-Natal, South Africa
Straw-coloured fruit bat (Eidolon helvum), female flying carrying pup on front. Lamin, Gambia
Goldcrest (Regulus regulus) in flight, portrait, Parainen Uto, Finland. September
Hen harrier (Circus cyaneus) juvenile male, in flight, Parainen Uto, Finland. September
Grey-headed flying-fox (Pteropus poliocephalus) in flight, with tongue out after licking water from its body
Eurasian hoopoe (Upupa epops) in flight with a grub in its beak, Pusztaszer reserve, Hungary. May
Namib desert dune ant (Camponotus detritus), queen on sand with visible wing scars on her thorax, Swakopmund, Namibia
Western reef egret (Egretta gularis) running through shallow water with wings open, chasing small fish, Kotu creek, The Gambia
Glasswing butterfly (Greta oto) resting on leaf. Captive
Sparrowhawk (Accipiter nisus) juvenile flying, Parainen Uto, Finland. September
Goshawk (Accipiter gentilis) flying in snow, Mainua Kajaani, Finland. March
Common tern (Sterna hirundo) catching a mayfly, Hjalstaviken, Uppland, Sweden. Nordic Nature Photo Competition 2021 Highly Commended - Nordic Bird Photo of the Year category
European bee-eater (Merops apiaster) two perched on branch, Lower Silesia, Poland
Black-headed gull (chroicocephalus ridibundus) diving into water, hunting fish / shellfish at sunrise. Baltic Sea, Poland. Sequence 1 of 2
Common buzzard (Buteo buteo) perched on stump with wings spread, Northamptonshire, UK, January
Bearded vulture (Gypaetus barbatus) flying, Ordesa National Park, Pyrenees, Spain
Taiwanese squeaking silkmoth (Rhodinia verecunda) controlled conditions
Campbell albatross (Thalassarche impavida), flying off Campbell Island (seen in background), January
Male Black grouse (Tetrao / Lyrurus tetrix) in flight. Kuusamo, Finland, April
Ruby-throated hummingbird (Archilochus colubris) male landing on Virginia creeper (Parthenocissus quinquefolia). Hill Country, Texas, USA
Great grey owl (Strix nebulosa) in flight in woodland during snowfall. Kuusamo. Finland. February
Northern harrier (Circus hudsonius), two juvenile females hunting. North Park, Colorado, USA. August
Apollo butterfly (Parnassius apollo) female laying eggs on branch. Parainen, Finland. July
RF - Small tortoiseshell butterfly (Aglais urticae) nectaring on Verbena (Verbena sp). Cornwall, England, UK. September. (This image may be licensed either as rights managed or royalty free.)
Saddle-billed stork (Ephippiorhynchus senegalensis) flapping wings after preening. Liuwa Plain National Park, Zambia
Taiwan blue magpie, (Urocissa caerulea) flying, in Taipeh, Taiwan, endemic species
Black-eared kite, (Milvus migrans lineatus), Bi Tan Bay, Taiwan
Mute swan (Cygnus olor) stretching its wings after preening on a misty morning. Valkenhorst Nature reserve, Valkenswaard, The Netherlands, May
European bee eaters (Merops apiaster) one taking off another perched, Kiskunsagi National Park, Pusztaszer, Hungary. May
