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Starr Puffer (Arothron stellatus), juvenile swimming. Lembeh Strait, North Sulawesi, Indonesia
Sumatran tiger (Panthera tigris sondaica). Captive, with digitally added leaf pattern
Reef manta rays (Manta alfredi) filter feeding in atoll passes and lagoons Hanifaru Bay, Raa Atoll, Maldives
Coyote (Canis latrans) walking through deep winter snow. Yellowstone National Park, Wyoming, USA. January
Goshawk (Accipiter gentilis) plucking a Willow Grouse (Lagopus lagopus) Liminka, Finland, March
Mother of pearl (Pleuroptya ruralis) moth nectaring on Ragwort (Jacobaea vulgaris) at night. Surrey, England, UK. July
Garden bumblebee (Bombus hortorum) queen nectaring on Comfrey (Symphytum Hidcote Pink ) flowers. In garden, Surrey, England, UK. March
Lined seahorses (Hippocampus erectus) amongst corals, The Bahamas
Sanderling (Calidris alba) flock foraging at sunset, Britanny, France, October
Red squirrel (Sciurus vulgaris) sitting on fallen tree in pine woodland, Glenfeshie, Cairngorms National Park, Scotland, UK, August
Gannet (Morus bassanus) fluffy chick with its parent. Shetland Islands, Scotland, UK, August
American bison (Bison bison) male walking through deep snow. Hayden Valley. Yellowstone National Park, Wyoming, USA. January
Wild Eurasian lynx (Lynx lynx) at night with city lights and sky glow behind, Switzerland. December. Taken with remote camera trap
Carp (Cyprinus carpio) in oxbow lake of the Aare River, Switzerland
Brown bear (Ursus arctos) portrait in the Ozernaya River, Kuril Lake, South Kamchatka Sanctuary, Russia
Robin (Erithacus rubecula) feeding on Gelder Rose berries, Germany. January, January
Coconut / Veined octopus (Amphioctopus marginatus) hunts in the sand at night, while illuminated by a divers torch beam, Anilao, Batangas, Luzon, Philippines. Verde Island Passages, Pacific Ocean
Zanzibar red colobus monkey (Procolobus kirkii) crossing the main road through the Reserve, Jozani forest, Jozani Chwaka Bay NP, Zanzibar, Tanzania, August
Marine iguana (Amblyrhynchus cristatus) on shore, Punta Moreno, Isabela Island, Galapagos
Bearded pig (Sus barbatus) digging in sand, foraging for crabs on beach, Bako National Park, Sarawak, Borneo
Grey whale (Eschrichtius robustus) tail, San Ignacio Lagoon, El Vizcaino Biosphere Reserve, Baja California, Mexico, March
Coal tit (Periparus ater) on mossy log with reflection, England, UK. January
Ladder-backed Woodpecker (Picoides scalaris), adult male perched on icy branch of Possum Haw Holly (Ilex decidua) with berries, Hill Country, Texas, USA. February
Violet-tailed sylph hummingbird (Aglaiocercus coelestis) Mindo, Pichincha, Ecuador
White prominent moth (Leucodonta bicoloria) County Kerry, Ireland
White ibis (Eudocimus albus) group of four in flight above clouds, Fort Myers Beach, Gulf Coast, Florida, USA, March
RF- Mute swan (Cygnus olor) on water in winter dawn mist. Loch Insh, Cairngorms National Park, Highlands, Scotland UK, December
Red squirrel (Sciurus vulgaris) gnawing red deer antler for minerals, Cairngorms National Park, Highlands, Scotland, UK, June
Red squirrels (Sciurus vulgaris) interacting, Cairngorms National Park, Highlands, Scotland, UK, January
Hen Harrier (Circus cyaneus) adult female in flight over heather moorland, Scotland, UK. July
King cobra (Ophiophagus hannah) in strike pose, Malaysia
Kamchatcka brown bear (Ursus arctos beringianus) with face underwater whilst fishing for Sockeye salmon in the Ozernaya River, Kuril Lake, South Kamtchatka Sanctuary, Russia, August
RF- Quarter horses running in snow at ranch, Shell, Wyoming, USA, February
Hoatzins (Opisthocomus hoazin) perched in tropical rainforest, Tambopata Reserve, Peru, South America
Tree sparrow (Passer montanus) feeding a fledgling on a branch of Hawthorn (Crataegus monogyna) blossom, Perthshire, Scotland. June 2013
Badger (Meles meles) cub amongst long grass, Dorset, England, UK, July
Red fox (Vulpes vulpes) resting in the snow, Churchill, Cananda, November
Whooper swans (Cygnus cygnus) two in flight, during snowfall, Lake Kussharo, Japan, February
Yellow-plumed parakeets (Leptosittaca branickii) at nest cavity in wax palm stump, Tapichalaca Biological Reserve, Ecuador
Barn owl on post in flooded Somerset Levels. England, UK, Europe
Bateleur eagle (Terathopius ecaudatus) head portrait, Captive, occurs in Africa
Portrait of a Common Wasp (Vespula vulgaris). UK. Focus stacked image (dead specimen)
Red squirrel (Sciurus vulgaris) having a drink, Black Isle, Scotland, UK. October
Long-eared owl (Asio otus) in a large tree, Cheshire, UK, March
Backlit photo of a Robust ghost pipefish (Solenostomus cyanopterus) reveals a small fish that it has just eaten in its stomach. Lembeh Strait, North Sulawesi, Indonesia. Molluca Sea
Wren (Troglodytes troglodytes) on frost covered fern, Dunham Massey, Cheshire, UK. Winner of the Habitat Category of the British Wildlife Photographer of the Year Awards (BWPA) 2017
Brandts cormorant (Phalacrocorax penicillatus) bursts through a school of Pacific chub mackerel (Scomber japonicus) while hunting beneath an oil rig
Corys shearwaters (Calonectris diomedea) diving among a mass of shoaling fish to feed, along with Atlantic spotted dolphins (Stenella frontalis), Formigas Islet dive site, Azores, Portugal