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Badger (Meles meles) adult, backlit at sunset, scratching its ears, Derbyshire, UK, May. Winner of the Animal Behaviour category of the British Wildlife Photography Awards (BWPA) Competition 2017
RF - Red Deer stag (Cervus elaphus) in snow-covered pine forest. Scotland, UK. December
Red deer stag (Cervus elaphus) in heavy snow, Cheshire, UK, March
Atlantic Puffin (Fratercula arctica) one landing among resting group, Sule Skerry, Scotland, UK, July
Sand lizard male {Lacerta agilis} Ainsdale NNR, Lancashire, UK
RF- Western clownfish (Amphiprion oceallaris) in Magnificent sea anemone (Heteractis magnifica). Raja Ampat, West Papua, Indonesia
Blue whale (Balaenoptera musculus) underwater, Coronado Islands, Baja California, Mexico, Eastern Pacific Ocean
Piraputanga (Brycon hilarii) shoal, Rio Olhio d agua, tributary of Rio da Prata, Bonito area, Serra da Bodoquena (Bodoquena Mountain Range), Mato Grosso del Sul, Brazil
Wolf (Canis lupus) pack running in deep snow, captive in enclosure of the Bavarian Forest National Park, Germany, February
Female Scottish wildcat (Felis silvestris grampia) portrait, in winter coat, semi habituated living wild in native oakwood, Lochaber, Scotland, April
Atlantic puffin (Fratercula arctica) gathering around burrows on cliff top, Isle of Lunga, Treshnish Isles, Isle of Mull, Scotland, UK, June
Adult Adder (Vipera berus) basking on pebble beach. Abbotsbury, Dorset, UK, March
Silhouette of Barn Swallow (Hirundo rustica) flying over water, hawking for insects, Berwickshire, Scotland, UK, July
Impala with Oxpeckers. Kruger National Park, South Africa
Emperor penguins (Aptenodytes forsteri) explode out of the water, returning to breed at Cape Washington, Antarctica. Taken on location for BBC Frozen Planet series, 2009
Squid (Abraliopsis atlantica) deep sea species from Atlantic Ocean off Cape Verde. Captive
Coastal brown bear (Ursus arctos) and cub, Lake Clarke National Park, Alaska, September
Black Bear (Ursus americanus) sleeping in a Sitka Spruce tree (Picea sitchensis) alongside Anan Creek, Alaska, July
Young Polar bear (Ursus maritimus) standing and trying to balance in shallow water along the Bernard Spit, 1002 area of the Arctic National Wildlife Refuge, North Slope of the Brooks Range, Alaska
Giant panda (Ailuropoda melanoleuca) cub yawning. Yuan Meng, first giant panda ever born in France, age 10 months, Captive at Beauval Zoo, Saint Aignan sur Cher, France
Polar bear cub 3 months (Ursus maritimus) playing in the front of the day den in march. Wapusk National Park. Churchill, Manitoba, Canada
Lion (Panthera leo) cubs playing, Masai-Mara Game Reserve, Kenya. Vulnerable species
Three Sperm whales (Physeter macrocephalus) socialising under the surface, Dominica, Caribbean Sea, Atlantic Ocean, Vulnerable species
RF- African elephant (Loxodonta africana) at sunset. Chobe National Park, Botswana. Endangered species
Trap-jaw ant (Odontomachus hastatus) with mandibles open, Los Amigos Biological Station, Peru
Mute swan (Cygnus olor) in early morning light removing its own feathers, during moulting, Valkenhorst Nature Reserve, The Netherlands, Europe. August
Goldfinch (Carduelis carduelis) feeding on teasel in falling snow, Leicestershire, UK, February
Calleta silkmoth (Eupackardia calleta), Texas USA Controlled conditions
Southern cassowary (Casuarius casuarius), adult male with chicks keeping a wary eye on the photographer. Near Bloomfield, north Queensland, Australia, September
Eyrean Grasswren (Amytornis goyderi) in typical hopping motion, Andado Station, Northern Territory, Australia, November
Bearded reedling (Panurus biarmicus) female feeding on reed seeds covered in frost, Espoo, Finland, January
Pletail unicornfish (Naso brevirostris) juvenile, its horn will continue to grow as it gets older. Fiji Pacific Ocean
Tabular iceberg floating in Weddell Sea, iceberg broken away from Larson C ice shelf. Adelie penguin (Pygoscelis adeliae) colony dwarfed on sea ice below. Near Danger Islands, Antarctica
White-tailed eagle (Haliaeetus albicilla) taking off with fish prey, Norway, August
Portrait of a Black-capped chickadee, (Poecile atricapillus) with white background, Block island, Rhode Island, USA. Bird caught during scientific research
Eurasian crag martin (Ptyonoprogne rupestris) Kawakarpo Mountain, Meri Snow Mountain National Park, Yunnan Province, China. May
Grey heron (Ardea cinerea) wading at night, reflected in water. Lincolnshire, UK, September
RF - White-legged damselfly (Platycnemis pennipes) peering over Oxeye daisy (Leucanthemum vulgare) flower. The Netherlands. August
Siberian tiger (Panthera tigris altaica) female territorial marking by rubbing cheek against ancient Manchurian fir tree, Land of the Leopard National Park, Far East Russia, November
Pair of Caribbean reef sharks (Carcharhinus perezi) swim over a coral reef. Jardines de la Reina, Gardens of the Queen National Park, Cuba. Caribbean Sea
Gaur (Bos gaurus) Manas National Park UNESCO World Heritage Site, Assam, India
Red fox (Vulpes vulpes) in snowy landscape with trees and two Crows (Corfus corone) winter, Jura, Switzerland
Green turtles (Chelonia mydas) swimming with angelfish, Punta Vicente Roca, Isabela Island, Galapagos
Great blue heron (Ardea herodias) taking off, with coastal landscape, Las Bachas, Santa Cruz Island, Galapagos
Wren (Troglodytes troglodytes) singing, early morning in spring, Broxwater, Cornwall, UK. April
Resplendent quetzal (Pharomachrus mocinno) male, Talamanca mountains, Costa Rica
Small Tortoiseshell Butterfly (Aglais urticae) feeding on scabious flowers in hay meadow, Wensum Valley, Norfolk, England, UK, July
Kiang (Equus kiang) herd grazing on steppe in evening light, mountains in background. Hoh Xil Nature Reserve, Tibetan plateau, Qinghai, China. October 2019