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Spotted jellyfish (Mastigias papua) Inanuran Island, Danajon Bank, Central Visayas, Philippines, April
African elephant (Loxodonta africana) herd walking in procession, Kaokoveld Desert, Namibia
Himalayan Blue sheep (Pseudois nayaur) Hemas National Park, Ladakh, India
Walia ibex (Capra walie) herd, Simien Mountains National Park, Ethiopia
Pig-nosed frog 1+Nasikabatrachus sahyadrensis+2 in leaf litter, Western Ghats, India
Japanese fireflies (Luciola cruciata) in flight at night, endemic species, Yaku-shima UNESCO World Heritage Site, Kagoshima, Japan, May
Wild Iberian Lynx (Lynx pardinus) male, Sierra de Andujar Natural Park, Jaen, Andalucia, Spain
Lion (Panthera leo) male smelling, flehmen response, Masai-Mara Game Reserve, Kenya. Vulnerable species
White rhino (Ceratotherium simum) female portrait, portrait, Nakuru National Park, Kenya
Female Pintail (Anas acuta) with chicks, Katmai National Park and Preserve, Alaska, USA, July
Red Foxes (Vulpes vulpes) young play fighting. Kronotsky Zapovednik Nature Reserve, Kamchatka Peninsula, Russian Far East, July
Male Rock ptarmigan (Lagopus mutus) vocalising whilst in flight, showing winter plumage, Utsjoki, Finland, April
Red Fox (Vulpes vulpes) curled up, with an eye open. Kronotsky Zapovednik Nature Reserve, Kamchatka Peninsula, Russian Far East, March
Gannet (Morus bassanus) adult turning in the air as it begins to plunge dive. Shetland Islands, Scotland, UK, July
Female Harpy Eagle (Harpia harpyja) perched close to its nest. Pousada Currupira d Araras, south west Brazil
Red-vented bulbul (Pycnonotus cafer) Bandhavgarh National Park, India, March
European Bison (Bison bonasus) calf and mother portrait. Mefody, first calf born to reintroduced herd in Bryansk Forest. Bryansk Forest Zapovednik, Kamchatka, Russian Far East, May
Great Frigatebird (Fregata minor) in flight over Midway Atoll. Midway
Baby Grey Squirrel (Sciurus carolinensis) kissing a tabby kitten
Great Blue shark (Prionace glauca) viewed from just below the surface, Pico Island, Azores, Portugal, Atlantic Ocean
European Brown Bear (Ursus arctos arctos), a large male taking a rest in a cloud of mosquitoes. Kajaani, Finland. June
Wood sandpiper (Tringa glareola) at dusk silhouetted against setting sun, Karigasniemi, Northern Finland, May. Bookplate from Danny Greens The Long Journey North
Himalayan marmot (Marmota himalayana) sun bathing on rock, Mount Kailash, Tibet
German Shepherd Dog bitch pup, Coco, 14 weeks old, with raised paw
West African / Niger Giraffe (Giraffa camelopardalis peralta) mother and baby. Zoo of Doue La Fontaine, France. Captive
Wilsons Storm Petrel (Oceanites oceanicus) feeding whilst walking on water Atlantic ocean, Madeira Portugal, August
Roseate Tern (Sterna dougallii) in flight in front of building, Madeira Portugal August
American Red Fox (Vulpes vulpes) cubs playing with its mother. Grand Teton National Park, USA, June
American Purple Gallinule (Porphyrio martinica) displaying brightly coloured wings. Everglades National Park, Florida, USA, March
Regal Jumping Spider (Phidippus regius) female. Captive, endemic to North America
Whale shark (Rhincodon typus) mouth open filter feeding at the surface, Quintana Roo, Yucatan Peninsular, Mexico, Caribbean Sea
Andean condor (Vultur gryphus) in flight over mountain peaks, Torres Del Paine National Park, Chile
Fallow deer (Dama dama) buck bellowing at dawn during the rut, Cheshire, UK, October
Common frog (Rana temporaria) leaping into a pond, controlled conditions, UK
Ural Owl (Stix uralensis) perched in snowy tree, Kuusamo Finland February
Atlantic Salmon (Salmo salar) in pool on upstream spawning migration, silhouetted against sky. Quebec, Canada, August
Wild Cat (Felis silvestris) in mist. Vosges, France, August
Tibetan antelope (Pantholops hodgsoni), two males, Kekexili, Qinghai, Tibetan Plateau, China, December
Lion cub (Panthera leo) silhouetted at sunrise, taken on location for Pride tv series. September 2003
Blue and white Border Collie puppy running forward
Two small pearl-bordered fritillary butterflies (Boloria selene) resting on bluebell, Marsland mouth, North Devon, UK. May 2017
Harvest mouse (Micromys minutus) on wheat stem, Devon, UK (captive). May
RF- Looking up under the gills of toadstools of Porcelain fungus (Oudemansiella mucida), Cornwall, UK
Five-spot burnet moths {Zygaena trifolii} mating on Pyramidal orchid {Anacamptis pyramidalis} Cornwall, UK
American red squirrel (Tamiasciurus hudsonicus) collecting, Douglas Fir (Pseudotsuga menziesii) cones, Montana, USA
White tent making bat (Ectophylla alba) roosting in tree, La Selva Field Station, Costa Rica
Great green macaw (Ara ambiguus) flying, La Selva Field Station, Costa Rica
Swift (Apus apus) in flight, Monmouthshire, Wales, UK, July