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Little blue penguin (Eudyptula minor), two standing on rocks at night, silhouetted against Melbourne city lights. St Kilda breakwater, Victoria, Australia. December 2016
Red squirrel (Sciurus vulgaris), two curled up asleep in drey inside nest box. Nest of lichen and Pine needles. Highlands, Scotland, UK. Medium repro only
Humpback whale (Megaptera novaeangliae) breaching, near Hout Bay, South Africa
Buzzard (Buteo buteo) in flight, portrait, Marlborough Downs, Wiltshire, UK. December
Brown bear (Ursus arctos) and Grey wolf (Canis lupus) together in wetlands, Kuhmo, Finland, July
Vietnamese Mossy Frog {Theloderma corticale} captive, from Vietnam
Common swifts (Apus apus) flying overhead, Wiltshire, UK, June. Digital composite image
Axolotl {Siredon / Ambystoma mexicanum} albino, captive
Starling murmuration, hundreds of thousands of starlings (Sturnus vulgaris) gather in the skies near their night time roost. Gretna Green, Scotland, January
Gemsbok (Oryx gazella) in sand dunes, Namibia. Finalist in the Mammals Category of the Wildlife Photographer of the Year Awards (WPOY) Competition 2015
African wild dogs (Lycaon pictus) juveniles playing with the leg of an impala, trying to drag it in three different directions. Mkuze, South Africa
Polar bear (Ursus maritimus) walking on Champ Island glacier above the sea, Franz Josef Land, Russian Arctic. Highly commended in Animals in their Environment Category of the Wildlife Photographer of
Tadpoles of the Common toad (Bufo bufo) swimming seen from below, Belgium, June
Axolotl (Ambystoma mexicanum), white or leucistic form, neotenic salamander. Captive, critically endangered in the wild, occurs in Mexico
Snowy owl (Bubo scandiaca) female, with lights behind, Canada, February
Blue whale (Balaenoptera musculus) diving beneath ocean surface. Indian Ocean, Sri Lanka
Shoebill stork (Balaeniceps rex) in the swamps of Mabamba, Lake Victoria, Uganda
Northern gannets (Morus bassanus) diving for fish, split level shot, Shetland, Scotland, UK, September
Polar bear (Ursus maritimus) on an iceberg, Baffin Bay, Canada. September
RF - Dapple grey Arabian stallion running in desert dunes near Dubai, United Arab Emirates
Arctic Fox (Vulpes lagopus), in winter coat portrait, Svalbard, Norway, April
Amazon horned frog (Ceratophrys cornuta) portrait, Yasuni National Park, Orellana, Ecuador
Aardvark (Orycteropus afer), young individual walking, Nambia. Captive, rescued individual
Killer Whales / Orcas (Orcinus orca)Killer whales / Orcas (Orcinus orca) large pod including calf traveling together while foraging on large schools of Herring (Clupea harengus) in the cold waters of northern Norway, January
Golden Eagle (Aquila chrysaetos) juvenile in flight, Norway, November
RF- Tawny owl with moon behind (Strix aluco), UK
RF - Hawksbill sea turtle (Eretmochelys imbricata) swimming over a coral reef. Whilst a Predatory bandcheek wrasse (Oxycheilinus digrammus)
Snowy owl (Bubo scandiaca) female in flight. Canada. February
Snow leopard (Panthera uncia) portrait with ears back. Captive
American bison (Bison bison) female covered in hoar frost near hot spring, portrait. Midway Geyser Basin, Yellowstone National Park, USA. February
Barn owl (Tyto alba) in flight, hunting, Hampshire, England, UK. Captive
Harpy Eagle (Harpia harpyja) portrait. Gamboa, Soberania National Park, Panama
Piraputanga fish (Brycon hilarii) in underwater landscape, Aquario Natural, Rio Baia Bonito, Bonito area, Serra da Bodoquena (Bodoquena Mountain Range), Mato Grosso do Sul, Brazil
Young Polar bear (Ursus maritimus) standing and trying to balance in shallow water along the Bernard Spit, 1002 area of the Arctic National Wildlife Refuge, North Slope of the Brooks Range, Alaska
Grizzly bear (Ursus arctos horribilis) female with cub riding on back, Katmai National Park, Alaska, USA, August
Leafy seadragon (Phycodurus eques) male carrying eggs, swims over seagrass meadow. Wool Bay, Edithburgh, South Australia. Gulf of St Vincent
Northern pike (Esox lucius) amongst Shining pondweed (Potamogeton lucens). Lake Neuchatel, Switzerland, September. Photographed for The Freshwater Project
Blue-footed booby (Sula nebouxii) courting pair, South coast, Santa Cruz Island, Galapagos
Tarpon (Megalops atlanticus) portrait, Eleuthera, Bahamas. December
Dugong (Dugong dugon) male feeding on a seagrass meadow (Halophila stipulacea), accompanied by a young Golden trevally (Gnathanodon speciosus). Marsa Nabaa, Marsa Alam, Egypt. Red Sea
Atlantic puffin (Fratercula arctica) landing, wings outstretched. Machias Seal Island, off the coast of Maine, USA. July
Eurasian bullfinch (Pyrrhula pyrrhula) male on a branch feeding on Rowan (Sorbus aucuparia) berries, Leon, Spain, February
Common loon (Gavia immer) with a chick alongside. British Columbia, Canada. June
Jay (Garrulus glandarius), two fighting in mid-air with another observing. Norway. October
Killer whale (Orcinus orca) male named Mel attacking young South American sea lion (Otaria flavescens) on beach, Punta Norte, Peninsula Valdes, Argentina
Herd of American bison (Bison bison) in snow, Yellowstone National Park, Wyoming, Yellowstone, January
Black and white view of Plains Zebra (Equus quagga) herd, Etosha National Park, Kunene, Namibia
Barn owl (Tyto alba) flying in snow covered countryside, Surrey, England, UK, January. Digital composite
