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Sunrise over Madison Mountains, Beaverhead National Forest, Montana, USA, July 2011
RF - Red fox (Vulpes vulpes) foraging in snow covered valley. Hayden Valley, Yellowstone National Park, USA. February 2019. (This image may be licensed either as rights managed or royalty free.)
Red fox (Vulpes vulpes) sequence of snow diving whilst hunting for rodents. Hayden Valley, Yellowstone National Park, USA. February 2019. Digital composite
North American river otter (Lutra canadiensis), two standing at edge of frozen Upper Yellowstone River, Hayden Valley, Yellowstone, USA. February
Steam rising from Grand Prismatic thermal pool in snow covered landscape. Midway Geyser Basin, Yellowstone National Park, USA. February 2019
American bison (Bison bison) female grazing near thermal pool, snow on ground. Biscuit Geyser Basin, Yellowstone National Park, USA. February 2019
Coyote (Canis latrans) standing near waters edge, in snow. Hayden Valley, Yellowstone National Park, USA. January 2019
Grizzly bear (Ursus arctos) running through water, Lake Clark National Park, Alaska, September
Soaptree yucca (Yucca elata) and twilight sky, showing Belt of Venus (blue of earth's shadow on the atsmophere)Soaptree yucca (Yucca elata) and twilight sky, showing Belt of Venus (blue of earth's shadow on the atsmophere below pink of sunset). White Sands National Monument, New Mexico, USA. November
Yellow mariposa lily (Calochortus luteus) with dark nectar guides. Cultivated in glasshouse, Surrey, England, UK. Endemic to California
Mesa Verdes Petroglyph Point, Mesa Verde National Park, Unesco World Heritage Site, Colorado, USA. October 2016
RF- American robin (Turdus migratorius) female feeding in Black hawthorn (Crataegus douglasii). Grand Teton National Park, Wyoming, USA
RF- Reindeer (Rangifer tarandus) feeding. Denali National Park, Alaska, USA
Brown bear Ursus arctos) searching for clams on tidal mudflat with waterbirds. Lake Clark National Park, Alaska. August
Hoodoo formation with caprock, at Stud Horse Point, Utah, January
Large leaves on Pacific blackberry (Rubus ursinus) above bracken fern and horsetail, is a small wet area. Oregon, USA. June
Sea otters (Enhydra lutris) resting on ice, Alaska, USA, June
Conifer sapling emerging from snow drift with cornice. Hayden Valley, Yellowstone National Park, Wyoming, USA. January
Wolf (Canis lupus) lone male walking in shadow of the Teton Mountains. Snake River valley near Jackson Hole. Grand Teton National Park, Wyoming, USA. January
Sun beam at sun rise over the Grand Canyon of the Yellowstone, Yelllowstone National Park, Wyoming, USA. January 2016
Coastal brown bears (Ursus arctos) fighting, Lake Clarke National Park, Alaska, September
Brandts cormorant (Phalacrocorax penicillatus) hunts for a meal in a school of Pacific chub mackerel (Scomber japonicus), beneath an oil rig
Young California sea lion (Zalophus californianus) play in the canopy of a forest of Giant kelp (Macrocystis pyrifera). Santa Barbara Island, Channel Islands. California, USA. North East Pacific Ocean
Coyote (Canis latrans) jumping in snow with its prey, Yellowstone National Park, Wyoming, USA, February
Two wild Mustang stallions fighting in White Mountain Herd Area, Wyoming, USA. August
Wild Mustang stallion chasing another away from his family in White Mountain Herd Area, Wyoming, USA. August
North Pacific right whale (Eubalaena japonica) swimming near surface, aerial view, Channel Islands National Park, California, USA. February
Arctic grayling (Thymallus arcticus) displaying his vibrant dorsal fin. North Park, Colorado, USA
Fort Collins- Colorado forest fire seen from Estes Park. The fire was intensified by dead trees killed by the Mountain pine beetle (Dendroctonus ponderosae) Colorado, USA, June
Pika (Ochotona princeps) gathering plants to store for winter, Bridger National Forest, Wyoming, USA. August
Sea otter (Enhydra lutris) pup sleeping, aged 6 days, floating, Monterey, California, USA
Sea otter (Enhydra lutris) mother and pup, aged 3 weeks, Monterey, California, USA
Red fox (Vulpes vulpes) two cubs playing, Shoshone National Forest, Wyoming, USA May
Least chipmunk (Tamias minimus) feeding on plant seeds, Grand Teton National Park, Wyoming, USA
North American Bison (Bison bison) coated in frost standing on snow, Yellowstone National Park, Wyoming, USA, January
Polar bear (Ursus maritimus) resting on sand, Kaktovik, Alaska, USA, September
Rotting wooden shed covered in moss, Washington State, USA
Humpback whale (Megaptera novaeangliae) blowing, Prince William Sound, Alaska, USA, July
American elk (Cervus elaphus canadensis) stag sticking tongue out, Yellowstone National Park, Wyoming, USA
RF- Caribou herd (Rangifer tarandus) grazing on tundra. Kobuk Valley National Park, Alaska, USA
Polar bear (Ursus maritimus) young bear in freezing water during autumn freeze up, Bernard Spit, 1002 area of the Arctic National Wildlife Refuge, North Slope, Alaska, Beaufort Sea
RF- Pinto Quarter horse in snow at ranch, Shell, Wyoming, USA, February
American barn swallow (Hirundo rustica erythrogaster) in flight catching insects over the Gallatin River, Yellowstone National Park, Montana, August
Saguaro cactus (Carnegiea gigantea) buds, Organ Pipe Cactus National Monument, Sonora Desert, Arizona, USA, April 2014
Rock formation caused by erosion in sandstone, Devils Garden, Grand Staircase-Escalante National Monument, Utah, USA, March 2014
American red squirrel (Tamiasciurus hudsonicus) collecting, Douglas Fir (Pseudotsuga menziesii) cones, Montana, USA. October
Yellow warbler (Dendroica petechia) collecting nesting material from Bulrush cattail (Typha sp) Bozeman, Montana. USA, June
Grizzly Bears (Ursus arctos) standing in heavy rain, Lake Clarke National Park, Alaska, September