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Hybrid Poplar tree plantation in autumn (Populs sp. ). Boardman Tree Farm, near IrrigonHybrid Poplar tree plantation in autumn (Populs sp.). Boardman Tree Farm, near Irrigon, Oregon, USA. October
North American beaver (Castor canadensis) on dam. Martinez, California, USA. June
American avocet (Recurvirostra americana), adult in breeding plumage foraging by sweeping its bill from side to side through shallow water, Orange County, California, USA, April
Clarks grebes (Aechmophorus clarkii) courting pair performing the Weed Ceremony in which they slowly circle breast to breast holding pond weed as nesting material, Lake Hodges, Escondido, California
Cinnamon teal (Anas cyanoptera), male taking flight from water, Orange County, California, USA, April
California Gull (Larus californicus), foraging for alkali flies (Ephydra hians) by running through swarm snapping its bill, on the shore at Mono Lake, California, USA. Digital frame expansion
California quail (Callipepla californica), male emerging from behind snowbank in winter, Mono Lake Basin, California, USA
Black-necked stilts (Himantopus mexicanus) pair performing post-copulatory display (as copulation ends, the male slides off the female leaving his wings draped over her)
Black-necked Stilt (Himantopus mexicanus), foraging in water, with reflection, Bear River Migratory Bird Refuge, Utah, USA, May
Red-breasted sapsucker (Sphyrapicus ruber) adult feeding on tree sap from shallow wells it has drilled in bark of Curl-leaf Mountain-mahogany (Cercocarpus ledifoius), Lee Vining Canyon, California
Crested auklets (Aethia cristatella) pair interacting while perched on rock, St Paul Island, Pribilofs, Alaska, USA, July
Horned puffins (Fratercula corniculata), one landing with outspread wings, to join its mate on cliff ledge, St. Paul Island, Pribilofs, Alaska, USA, July
Sea otter (Enhydra lutris) group of four resting among sea ice, Alaska, USA, June
Humpback Whale (Megaptera novaeangliae) blowing or spouting and fluking, Southeast Alaska, USA, August
Male American Bison (Bison bison) walking through snow. Firehole River Valley, Yellowstone National Park, Wyoming, USA. January
Trumpeter swans (Cygnus buccinator) on the edge of the Upper Yellowstone River. Hayden Valley, Yellowstone National Park, Wyoming, USA. January
Burrowing owl (Athene cunicularia) doing rain dance, spreading wings and dancing while it rained, Arizona, USA
Burrowing owl (Athene cunicularia) rotating head, Arizona, USA
Blacksmiths (Chromis punctipinnis) school swam through a giant kelp (Macrocystis pyrifera) forest, Santa Barbara Island, Channel Islands. Los Angeles, California, USA, North East Pacific Ocean
Giant sequoia (Sequoiadendron giganteum) trees in Sequoia National Park, California, USA, September 2014
Young Polar bear (Ursus maritimus) standing on hing legs, Bernard Spit, 1002 Area, Arctic National Wildlife Refuge, North Slope, Alaska, USA, October. Vulnerable species
Irregular ripples on gypsum sand dunes created by high winds, White Sands National Monument, Chihuahuan Desert, New Mexico, USA, December 2012
Polar bear (Ursus maritimus) sow with a pair of cubs walk on a barrier island during autumn freeze up, Bernard Spit, North Slope, Arctic coast of Alaska, September
Polar bear (Ursus maritimus) swimming at surface of Beaufort Sea, Alaska, USA
Polar bear (Ursus maritimus) sow standing along snow covered Bernard Spit, off the 1002 Area, Arctic National Wildlife Refuge, North Slope, Alaska, USA, October. Vulnerable species
Polar bear (Ursus maritimus) mother with juveniles resting on newly formed pack ice during autumn freeze up, Beaufort Sea, off Arctic coast, Alaska
Polar bear (Ursus maritimus) juvenile crossing over newly forming pack ice, during autumn freeze up, Beaufort Sea, off Arctic coast, Alaska
Quarter horses running in snow at ranch, Shell, Wyoming, USA, February
RF - Two Gypsy vanner geldings standing together looking away, Greyrock, Wyoming, USA. June
Two wild pinto Mustang stallions battle for dominance in Sand Wash Basin, Colorado, USA
Two wild Mustang stallions fighting in the White Mountain Herd Area, Wyoming, USA. August
Two Gypsy vanner geldings in wildflower pastures, Wyoming, USA. June 2014
Wild Mustang horse rearing, Pryor mountains, Montana, USA. June
RF - Head portrait of grey Andalusian mare with long mane in snow, Berthoud, Colorado, USA. January
Wild Mustang, dun horse in dust, Sand Wash Basin Herd Area, Colorado, USA
Texas Longhorn cow on high country pasture, Santa Ynez Mountains foothills, Goleta, California, USA
Wild red roan / pinto Mustang mare with bald faced foal in Adobe Town Herd Area, Wyoming, USAWild red roan/pinto Mustang mare with bald faced foal in Adobe Town Herd Area, Wyoming, USA. April
Wild horses, Monument valley tribal park, Navajo reserve, Utah, USA. April
Polar bear (Ursus maritimus), female with two cubs, on a barrier island outside Kaktovik, Alaska, USA
Wild Mustang horses interacting, Pryor mountains, Montana, USA. June
Polar bear (Ursus maritimus) silhouetted against setting sun, Bernard Spit, off the 1002 Area, Arctic National Wildlife Refuge, North Slope, Alaska, USA, October. Vulnerable species
Arctic ground squirrel (Spermophilus parryii) Katmai National Park, Alaska, USA, August
Humpback whale (Megaptera novaeangliae) breaching, Prince William Sound, Alaska, July
North American Bobcat (Lynx rufus) stalking along Madison River. Yellowstone National Park, Wyoming, USA. January
Rocky Mountain Goat (Oreamnos americanus) male on rocks at 14, 000 feet elevation, Mount Evans, Colorado, USA. July
Pronghorn (Antilocapra americana) male, Yellowstone National Park, Wyoming, USA, June
Group of California sea lions (Zalophus californianus) in a kelp forest (Macrocystis pyrifera), California, USA. Pacific ocean
Cinnamon bear, subspecies of black bear (Ursus americanus cinnamomum) cub climbing tree, Yellowstone National Park, Wyoming, USA, May
