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Gila woodpecker (Melanerpes uropygialis) male feeding on nectar in Saguaro cactus blossom (Carnegiea gigantea), Lost Dutchman State Park, Arizona, USA, April 2013
Northern hawk owl (Surnia ulula) perched, Nome, Alaska, USA, March
Snowy owl (Bubo scandiacus) on beach. Ocean County, Washington, USA. March
Ruddy turnstone (Arenaria interpres) in breeding plumage. Yukon Delta National Wildlife Refuge. Alaska, USA. June
Group of California sea lions (Zalophus californianus) swimming in kelp forest (Macrocystis pyrifera), California, USA. Pacific ocean
Quarter horses running in snow at ranch, Shell, Wyoming, USA, February
Skewbald Horse in ranch, Martinsdale, Montana, USA
Rocky mountain horse, Bozeman, Montana, USA. June
Prairie Rattlesnake (Crotalus viridis) sunbathing, Bozeman, Montana, USA
Tree swallow (Tachycineta bicolor) in fight over the Madison River, Montana, USA, June
American barn swallow (Hirundo rustica erythrogaster) in flight catching insects over the Gallatin River, Yellowstone National Park, Montana, August
Tree swallows (Tachycineta bicolor), perched on wire, calling aggressively to each other, Madison River, Montana, USA, June
Bald eagle (Haliaeetus leucocephalus) head portrait, Alaska, USA, February
Bighorn sheep (Ovis canadensis) close up rams head shot, Yellowstone National Park, USA, February
Polar bear (Ursus maritimus) portrait, female. Bernard Spit, 1002 area of the Arctic National Wildlife Refuge, North Slope, Alaska
Pink lady mayfly (Epeorus albertae) close up of eyes of male, Madison River, Montana, USA, August
Detail of the gas bladders and fronds of Giant kelp (Macrocystis pyrifera) plant. Santa Barbara Island, the Channel Islands, California, United States of America. East Pacific Ocean
Lodgepole pine cones (Pinus contorta), Teton County, Wyoming, USA. May
RF - Sea otter (Enhydra lutris) floating, California, USA, July
Black-bellied Plover (Pluvialis squatarola) chicks in the nest, seen from overhead, Yukon Delta National Wildlife Refuge, Alaska, USA June
Giant sequoia (Sequoiadendron giganteum) trees in Sequoia National Park, California, USA, September 2014
Canyon live oak (Quercus chrysolepis) Diablo Range, California, USA, June
Low angle view of trees in the Hoh Rainforest, Hall of Mosses Trail, Olympic National Park, Jefferson County, Washington, USA, June
Trunks of giant sequoia trees (Sequoiadendron giganteum) Sequoia National Park, California, USA, November 2012
Grand Teton peak, Grand Teton National Park, Wyoming, USA
RF - American bisons (Bison bison) in Grand Teton National Park. winter. Wyoming, USA
California sea lions (Zalophus californianus) two resting with flippers round each other, Monterey Bay, California, USA, September
Polar bear (Ursus maritimus) juvenile rolling around on newly formed pack ice, Beaufort Sea, off the 1002 area of the Arctic National Wildlife Refuge, North Slope, Alaska
Giant kelp (Macrocystis pyrifera) reaching up toward the sea surface, off Santa Barbara Island, California, USA, Pacific Ocean
Grizzly bear (Ursus arctos) female and nuzzling cub, Lake Clark National Park, Silver Salmon, Alaska. July
Sea otter (Enhydra lutris) female and pup playing. Alaska, USA, February
Panoramic of Point Lobos State Natural Reserve, Big Sur Coast, California, USA, September 2014
Bald eagle (Haliaeetus leucocephalus) flying over water in falling snow. Kachemak Bay, Alaska, USA. February
Rainclouds over Madison Valley, Beaverhead National Forest, Greater Yellowstone Ecosystem, Madision Mountains, Montana USA, July
Antelope Canyon, a slot canyon formed of Navajo sandstone and shaped by water erosion, Arizona, USA
Female American buffalo (Bison bison) walking across grassland with newborn calf, Yellowstone National Park, Wyoming, USA. May
Coyote (Canis latrans) hunting after a fresh snowfall, Yellowstone National Park, Wyoming, USA. October
Air bladders lifting strands of giant kelp (Macrocystis pyrifera) towards the surface, off Santa Barbara Island, California, USA
Annas hummingbird (Calypte anna) male dancing in flight and displaying his bright pink throat feathers to a female, Montana, USA. June
Barn and wheat field in the Palouse farming area of southeastern Washington, USA, August
Moose (Alces alces) bull crossing mountain river at sunset, Grand Teton National Park, Wyoming, USA
Coyote (Canis latrans) pouncing, hunting technique in Yellowstone National Park, Wyoming, USA
Great horned owls (Bubo virginianus), adult and chick, on nest in Saguaro cactus (Carnegiea gigantea), Sonoran desert, Arizona
Kelp bass (Paralabrax clathratus) in Giant kelp (Macrocystis pyrifera) forest, Santa Barbara, California, USA
Great horned owl (Bubo virginianus) adult and chick in nest in Saguaro cactus (Carnegiea gigantea) Sonoran Desert, Arizona, USA
Black-footed ferret (Mustela nigripes) running in the snow as it goes from one prairie dog burrow to another. Colorado, USA, Januay. January
Swift fox (Vulpes velox) female, portrait. Eastern Colorado, USA, May