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Southern stingray (Dasyatis americana) large female foraging over seagrass in shallow water. The Sandbar, Grand Cayman, Cayman Islands. British West Indies. Caribbean Sea
Coral Rabbitfish (Siganus corallinus) portrait, Tubbataha Reef Natural Park, UNESCO World Heritage Site, Sulu Sea, Cagayancillo, Palawan, Philippines
Crowned dwarf caiman (Paleosuchus trigonatus) male resting on a log. Yasuni National Park, Orellana, Ecuador
Cichlids (Cichlidae) in Lake Malawi, Malawi, November. Photographed for The Freshwater Project
Aquatic plants in a spring of the Gacka River, Croatia. October 2012. Photographed for The Freshwater Project
Ross Ice Shelf, the largest ice shelf of Antarctica, near Cape Crozier, Ross Island, Ross Sea, Antarctica Aerial shot from helicopter 02.02.2017 Trip on the MV Ortelius, 13 Jan
RF - Coscoroba swan (Coscoroba coscoroba) pair with chicks on water La Pampa, Argentina
Flower tree soft corals (Umbellulifera sp. ), Triton Bay, near Kaimana, West PapuaFlower tree soft corals (Umbellulifera sp.), Triton Bay, near Kaimana, West Papua, Indonesia
Striped mackerel (Rastrelliger kanagurta) shoal swimming with open mouths, filtering for zooplankton. Marsa Shouna, Port Ghalib, Marsa Alam, Egypt. Red Sea
Bioluminescent Sea-fireflies (Vargula hilgendorfii) producing a bright blue light. The light is produced by mixing two chemicals together in the presence of oxygen
Bluespine unicornfish (Naso unicornis) Shark Reef to Jolande, Ras Mohammed National Park, Egypt, Red Sea
Trumpetfish (Aulostomus maculatus) using a Spanish Hogfish (Bodianus rufus) as a stalking horse. Spanish Hogfish is eating an arm of a starfish. Bonaire, Leeward Antilles, Caribbean
Blue shark (Prionace glauca) off Halifax, Nova Scotia, Canada. July
Veined octopus (Amphioctopus marginatus) between clam shell halves. Ambon, Maluku Archipelago, Indonesia. Banda Sea, tropical west Pacific Ocean
Blue whale (Balaenoptera musculus) diving beneath the surface. Indian Ocean, Sri Lanka
Tawny nurse shark (Nebrius ferrugineus) group, individuals circling each other at dusk. Vavuu Atoll, Maldives. Indian Ocean
Pheasant (Phasianus colchicus) male calling, East Frisian Islands, Wittbulten National Park. Wadden Sea UNESCO World Heritage Site, Germany, June
Brydes whale (Balaenoptera edeni) Trincomalee, Eastern Province, Sri Lanka, Bay of Bengal, Indian Ocean
Pearlfish (Carapus acus) in anus of Sea cucumber (Stichopus regalis). These species have a commensal relationship, with the fish living in the gut of the sea cucumber. Catalonia, Spain
Sea butterfly (Peraclis bispinosa) captive deep sea species from Atlantic Ocean off Cape Verde
Glass octopus (Vitreledonella richardi). deep sea species from Atlantic Ocean off Cape Verde. Captive
Fangtooth moray eel (Enchelycore anatina) with mouth open, Grand Canaria, Canary Islands, Spain. East Atlantic Ocean
Humboldt squid (Dosidicus gigas) cannibalising another squid of the same species at night off Loreto, Sea of Cortez, Baja California, Mexico, East Pacific Ocean
Illuminated netdevil (Linophryne arborifera), female specimen, from deepsea Atlantic Ocean near Cape Verde at a depth of 1900m
Female Polar bear (Ursus maritimus) curiously checks out the photographer, standing upright on hind legs, Svalbard, Norway
Green Sea Turtle (Chelonia mydas) returning to sea after Egg laying, Raine Island, Australia. Raine Island is the largest and most important green sea turtle nesting area in the world, with over 14
Aggregation of thousands of Spider crabs (Leptomithrax gaimardii) for moulting, South Australia Basin, Australia. Pacific Ocean
Australian Black Swan (Cygnus atratus) Bay of Shoals, Kingscote, Kangaroo Island, South Australia State, Australia
Dwarf Minke whale (Balaenoptera acutorostrata) Great Barrier Reef, Australia. Coral Sea
Humpback whale (Megaptera novaeangliae) breaching, Prince William Sound, Alaska, July
Florida manatee (Trichechus manatus latirostrus), Three Sisters Springs, Crystal River, Florida, United States
Group of California sea lions (Zalophus californianus) in a kelp forest (Macrocystis pyrifera), California, USA. Pacific ocean
Grizzly bear (Ursus arctos horribils) sow sits in riverbed with a mountain range in background, east coast of Katmai National Park on the Alaskan peninsula, USA
RF- Florida manatee (Trichechus manatus latirostrus) portrait in Three Sisters Spring. Crystal River, Florida, USA. February 2010
Canada goose (Branta canadensis) adult swimming leading five goslings, Ithaca, New York, USA
Tisza mayfly (Palingenia Longicauda) in the Tisza river, Hungary, June 2009
RF- Mawddach Estuary at low tide, Barmouth, Snowdonia National Park, Gwynedd, Wales, May 2012
RF- Bannded demoiselle (Calopteryx splendens), resting on buttercup, Lower Tamar Lakes, Cornwall/Devon border, UK. May
RF- River Plym flowing through Dewerstone Wood, Dartmoor National Park, Devon, England, UK, October
A misty morning view looking down the River Tay in autumn, Kinnoull Hill Woodland Park, Perthshire, Scotland, November 2011
Dipper (Cinclus cinclus) on rock in stream. Perthshire, Scotland, May
View over New Forest heathland with Ling (Calluna vulgaris), Bell Heather (Erica cinerea) and pool. Fritham Cross, New Forest National Park, Hampshire, England, UK, August
Female Emperor dragonfly (Anax imperator) laying eggs, Cornwall, England, UK, April
Adult Daubentons bat (Myotis daubentoni) flying over a weir, England, UK, September. 2020VISION Exhibition
European river otter (Lutra lutra) climbing to the top of a weir, river, Dorset, UK, November
Avocet (Recurvirostra avosetta) trying to drive Shelduck (Tadorna tadorna) pair away from breeding grounds, Oare marshes, RSPB Greater Thames Futurescapes Project, North Kent, UK, June
The Cuckmere River at sunset, Seven Sisters Country Park, South Downs National Park, East Sussex, England, UK, July 2011
RF- Rear view of Basking shark (Cetorhinus maximus) feeding on plankton, visible as white dots on water surface near island of Coll, Inner Hebrides, Scotland, UK. June
