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Red deer (Cervus elaphus) stag bellowing, rutting season, Roehampton Flats in background, Richmond Park, London, UK, September
Red deer (Cervus elaphus) stag portrait, bellowing with bracken in antlers, rutting season, Bushy Park, London, UK, October
Firecrest (Regulus ignicapillus) male singing, Norfolk, UK, April. Did you know? Along with the goldcrest, the firecrest is Britains smallest bird
Sedge warbler (Acrocephalus schoenobaenus) adult singing in reedbed, Cambridgeshire, UK, May
Common nightingale (Luscinia megarhynchos) adult perched, singing, Cambridgeshire, UK, April. Photographer quote: Nightingales are more renowned for the beauty of their song than their looks
Red deer (Cervus elaphus) stag bellowing, Bradgate Park, Leicestershire, England, UK, October
Close up of Capercaillie (Tetrao urogallus) male displaying in pine forest, Cairngorms NP, Highlands, Scotland, UK, February
Red-crowned Cranes (Grus japonensis) displaying and calling in snow, Hokkaido, Japan, February
Brown Duckling cheeping, against white background
Common / Eurasian crane (Grus grus) calling at sunset, silhouetted. Lake Hornborga, Sweden, April
RF- Grey Wolf (Canis lupus) howling at sunset (digitally enhanced)
Razorbill (Alca torda) pair calling, between soft focus vegetation. Great Saltee, Saltee Islands, Co. Wexford, Ireland, June
Whooper Swans (Cygnus cygnus) on snow, Caerlaverock WWT, Scotland, Solway, UK, January
Male Masked trogon (Trogon personatus) portrait, Mindo, Pichincha, Ecuador
Andean Gull (Chroicocephalus serranus) calling in flight, Cotopaxi, Ecuador
Sedge warbler (Acrocephalus schoenobaenus) singing portrait, Saltee Islands, Republic of Ireland, May. Non-ex
Capercaillie (Tetrao urogallus) male displaying in pine forest in snow, Cairngorms NP, Highlands, Scotland, UK, January. Photographer quote
Great skua (Stercocarius skua) in flight over Gannet (Morus bassanus) colony, looking for small chicks, whilst Gannets call defensively, Shetland Isles, July
Two Gannets (Morus bassanus) squabbling, watched by another in the background. Hermaness National Nature reserve, Shetland Isles, Scotland
Capercaillie (Tetrao urogallus) male, displaying in pine forest, Cairngorms NP, Scotland, UK, April
European brown tabby kitten, vocalising, Italy
Eagle owl portrait {Bubo bubo} captive, Germany
Snow goose with gosling (Chen caerulescens) at sunset. Taken on location for BBC Frozen Planet series, 2008
Arctic tern (Sterna paradisaea) adult hovering directly in front of the sun, defending nest from perceived threat, Farne Islands, UK. June Non-ex
Couchs Spadefoot (Scaphiopus couchii) adult at night calling with vocal sac inflated. Laredo, Webb County, South Texas, USA, April
Laysan albatross (Phoebastria immutabilis) head portrait calling mate, Guadalupe Island Biosphere Reserve, off the coast of Baja California, Mexico, March
Arctic tern (Sterna paradisaea) in flight, vocalising, aggresive behaviour, Farne Islands, Northumberland, UK, June
Arctic tern (Sterna paradisaea) in flight, calling, Farne Islands, Northumberland, UK, June
Common cranes {Grus grus} calling with heads raised in air, Laguna de Gallocanta, Teruel, Aragn, Spain
Corncrake (Crex crex) calling from rock, North Uist, Outer Hebrides, Scotland, UK
RF - Chinstrap Penguins (Pygoscelis antarcticus) once calling, South Shetland Islands, Antarctica Continent
Gentoo penguins on iceberg (Pygoscelis papua) Antarctic Peninsula
Alpine marmot (Marmota marmota) calling, Hohe Tauern National Park, Austria, July
Lynx (Lynx lynx) cub calling. Hannau, Germany, September. Captive
