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Spotted hyaena laughing {Crocuta crocuta} Kenya
Red deer (Cervus elaphus) stag bellowing during rut, Derbyshire, UK
Common Raven (Corvus corax) two in flight during snow storm, Kuusamo, Finland, April
Bluethroat (Luscinia svecica) male singing, Sinirinta, Kuusamo, Finland, June. Fascinating birds bookplate
Asiatic wild ass (Equus hemionus) with foal, captive, UK, endangered species
Three Brown-hooded Parrots (Pyrilia haematotis) one taking off from branch, in the rain Laguna del Lagarto, Santa Rita, Costa Rica
Yellow-headed Blackbird (Xanthocephalus xanthocephalus) male calling and performing courtship / territorial display in cattail marsh, Mono Lake Basin, California, USA
Red howler monkey (Alouatta seniculus) howling, captive
Black backed jackal {Canis mesomelas} howling, Etosha NP, Namibia
Two male Ostriches (Struthio camelus) running during dispute, Etosha NP, Namibia
Gentoo penguins (Pygoscelis papua) calling, Cuverville Island. Antarctic Peninsula, Antarctica
Common yellow throat (Geothlypis trichas) male, singing, Anchorage Provincial Park, Grand Manan Island, New Brunswick, Canada, June
Robin (Erithacus rubecula) singing on bare tree branch, Whitlingham Country Park, Norfolk, UK. February
Yellow-bellied toad (Bombina variegata) male in pond, calling, Lucerne, Switzerland. July
Bluethroat (Luscinia svecica cyanecula) male, perched on tree stump singing, Germany. March
Little bittern (Ixobrychus minutus) perched in reedbed calling, Valkenhorst nature reserve, Valkenswaard, the Netherlands. May
Robin (Erithacus rubecula) perched on branch singing in spring, Bavaria, Germany. April
Bluethroat (Luscinia svecica cyanecula) male, singing as a courtship display, Germany
Ragdoll-cross kitten, aged 3 weeks, meowing, portrait
Galapagos fur seal (Arctocephalus galapagoensis) with mouth open, head portrait, Santiago Island, Galapagos National Park, Galapagos Islands. Endangered
Fosa (Cryptoprocta ferox) pair, mating in forest canopy, Zombitse-Vohibasia National Park, central south western Madagascar
Black Welsh mountain sheep lamb calling. Wales UK, March
Bengal tiger (Panthera tigris tigris) walking through river, snarling, Bardia National Park, Terai, Nepal. Endangered
Green bee-eater (Merops orientalis) calling, portrait, Bardia National Park, Terai, Nepal
Three Brown noddies / Common noddies (Anous stolidus) trying to steal a fish from a Brown pelican (Pelecanus occidentalis), Santiago Island, Galapagos Islands, Pacific Ocean
Northern parula (Parula americana) male singing from flowering crabapple (Malus sp)Northern parula (Parula americana) male singing from flowering crabapple (Malus sp.) tree in spring, near Salamanca, New York, USA, May 2022
Common pheasant (Phasianus colchicus) male flapping wings and calling, Pusztaszer reserve, Hungary. May
Male Western capercaillie (Tetrao urogallus) calling during lek mating, Finland. March
Pair of adult Red-necked grebe (Podiceps grisegena) calling, Finland. May
Jackdaw (Corvus monedula) swooping down calling with claws raised to threaten a Grey squirrel (Sciurus carolinensis) in the entrance to a nest box the bird wants to nest in which is already occupied
Red deer (Cervus elaphus) stag bellowing amongst bracken, Bushy Park, London, England, UK, October
Robin (Erithacus rubecula) perching on a branch, singing, Bavaria, Germany. May
Dominant male Chimpanzee (Pan troglodytes troglodytes) in the mangrove, with female and infant vocalising, Conkouati-Douli National Park, Republic of Congo, Africa
Spoonbill (Platalea leucorodia) calling in shallow pond on misty morning, Valkenhorst Nature Reserve, The Netherlands, August
Sedge warbler (Acrocephalus schoenobaenus) singing with bokeh effect in background, Norfolk, England, UK, May. Winner of the Birds in Environment Category of the Bird Photographer of the Year 2015
Sub-Antarctic / Brown skua (Stercorarius antarcticus) flying away with egg stolen from a Royal penguin colony (Eudyptes schlegeli), Macquarie Island, Australian Territory. December
Southern elephant seal (Mirounga leonina) male, rests and yawns on the snow. Right Whale Bay, South Georgia Island
Northern spadefoot (Notaden melanoscaphus) adult male with vocal sac inflated, calling, Howard River sand flats, Northern Territory, Australia, January
Bilingual frog (Crinia bilingua) adult male calling, Adelaide River, Northern Territory, Australia, January
Saddleback anemonefish (Amphiprion polymnus) barks a warning as it guards a clutch of developing eggs (beneath it) laid on a small stone, below a fold in its home
Red deer (Cervus elaphus) in the pine forest bellowing during the rutting season. Barycz Valley Landscape Park, Poland. September
Robin (Erithacus rubecula) male singing, perched on branch. Joutsa, Finland. April
Short-headed treefrog (Dendropsophus brevifrons) calling with inflated vocal sac. Above a rainforest pond at night, Yasuni National Park, Ecuador
Ring-tailed lemur (Lemur catta) suckling baby. Berenty Reserve, Madagascar
European robin (Erithacus rubecula) singing. London. January
Black-bellied bustard (Lissotis melanogaster) male calling at sunrise. Serengeti National Park, Tanzania
African grey hornbill (Lophoceros nasutus) calling in display, perched on tree snag. Janjanbureh, Gambia