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Gyrfalcon (Falco rusticolus) juvenile calling, Thingeyjarsyslur, Iceland, July 2009
Two White stork (Ciconia Ciconia) chicks, one calling, Prypiat area, Turov, Belarus, June 2009
Red deer (Cervus elaphus) stag calling at sunset during rut, Klampenborg Dyrehaven, Denmark, September 2008
Alpine marmot (Marmota marmota) giving an alarm call, Guillestre, Queyras, France, June 2009
White stork (Ciconia ciconia) in long grass calling, Kurmene, Latvia, June 2009
Great snipe (Gallinago media) calling, Matsalu National Park, Estonia, May 2009
Great snipe (Gallinago media) shaking head calling, Matsalu National Park, Estonia, May 2009
Marsh warbler (Acrocephalus palustris) male singing at dawn, Lithuania, June 2009
Marsh frog (Rana ridibunda) calling, Sierra de Andjar Natural Park, Mediterranean woodland of Sierra Morena, north east Jan Province, Andalusia, Spain, April 2009
Whinchat (Saxicola saxicola) male singing perched on post, Lithuania, May 2009
White stork (Ciconia ciconia) adult in breeding plumage on nest, calling, Lithuania, May 2009
Two Steppe eagle (Aquila nipalensis) chicks, Cherniye Zemli (Black Earth) Nature Reserve, Kalmykia, Russia, May 2009
European souslik (Spermophilus citellus) feeding, Eastern Slovakia, Europe, June 2009
Great reed warbler (Acrocephalus arundinaceus) perched on reed, singing, Eastern Slowakia, Europe, June 2009
Black stork (Ciconia nigra) at nest calling, Eastern Slovakia, Europe, June 2009
Avocet (Recurvirostra avosetta) calling, Texel, Netherlands, May 2009
Two Common terns (Sterna hirundo) one with fish in its beak, the other calling, Texel, Netherlands, May 2009
Tree sparrow (Passer montanus) displaying in rose bush, Slovakia, Europe, May 2009
Black tailed godwit (Limosa limosa) standing on one leg on post calling, Texel, Netherlands, May 2009
Capercaillie (Tetrao urogallus) male calling as part of display, Cairngorms NP, Scotland, February 2009. WWE INDOOR EXHIBITION
Capercaillie (Tetrao urogallus) male displaying in snow, Cairngorms NP, Scotland, February 2009
Red deer (Cervus elaphus) stag under trees calling during rut in morning, Klampenborg Dyrehaven, Denmark, September 2008
Red deer (Cervus elaphus) stag calling during rut, light mist at sunrise, Klampenborg Dyrehaven, Denmark, September 2008. WWE INDOOR EXHIBITION
Red deer (Cervus elaphus) calling at sunset, during rut, Klampenborg Dyrehaven, Denmark, September 2008. WWE INDOOR EXHIBITION
Grey heron (Ardea cinerea) with beak open, Elbe Biosphere Reserve, Lower Saxony, Germany, September 2008
Red deer (Cervus elaphus) stag calling during rut, Klampenborg Dyrehaven, Denmark, September 2008. WWE BOOK. WWE OUTDOOR EXHIBITION. NOT AVAILABLE FOR GREETING CARDS OR CALENDARS *PRESS IMAGE
Puffin (Fratercula arctica) standing on rock calling, Saltee Islands, County Wexford, Ireland, June 2009
Corncrake (Crex crex) calling, Tory Island, County Donegal, Ireland, June 2009
Razorbill (Alca torda) calling, Saltee Islands, County Wexford, Ireland, June 2009
Shag (Phalacrocorax aristotelis) calling with wings stretched out, Saltee Islands, County Wexford, Ireland, June 2009
Red throated diver (Gavia stellata) calling, in breeding plumage, Bergslagen, Sweden, April 2009
Little auk (Alle alle) calling, Fuglesangen, Svalbard, Norway, June 2008
Cyprus pied wheatear (Oenanthe cypriaca) perched on barbed wire singing, Northern Cyprus, April 2009
Eleonoras falcon (Falco eleonorae) taking off from rock ledge, Andros, Greece, September 2008. WWE BOOK. WWE INDOOR EXHIBITION
Grey seal (Halichoerus grypus) mating behaviour, Donna Nook, Lincolnshire, UK, November 2008
Grey seal (Halichoerus grypus) pup calling, Donna Nook, Lincolnshire, UK, November 2008
Grey seal (Halichoerus grypus) pair calling, Donna Nook, Lincolnshire, UK, November 2008
European Bee-eater (Merops apiaster) vocalising, Pusztaszer, Hungary, May 2008
European green toad (Bufo viridis) calling, vocal sac inflated, with eyes closed, in a pond (at 2, 711m) Adylsu valley, side valley to Baksan valley and Elbrus, Caucasus, Russia, June 2008
Skylark (Alauda arvensis) singing on branch stump, East Slovakia, Europe, June 2008
European souslik (Spermophilus citellus) calling, Slovakia, Europe, June 2008
European Roller (Coracias garrulus) calling, Pusztaszer, Hungary, May 2008
Alpine marmot (Marmota marmota) aggressive behaviour, Hohe Tauern National Park, Austria, July 2008
Alpine marmot (Marmota marmota) alert, Hohe Tauern National Park, Austria, July 2008
Black-headed bunting (Emberiza melanocephala) male perched singing, Bulgaria, May 2008
Golden oriole (Oriolus oriolus) pair at nest, Bulgaria, May 2008
Red deer (Cervus elaphus) stag bellowing, rutting season, Roehampton Flats in background, Richmond Park, London, UK, September
Red deer (Cervus elaphus) stag portrait, bellowing with bracken in antlers, rutting season, Bushy Park, London, UK, October