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Desert chameleon {Chamaeleo namaquensis} note one eye facing forwards, the other backwards, Namib desert, Namibia
Lioness in long grass {Panthera leo} Linyanti, Botswana
Two female Leaf nosed twig snakes {Langaha madagascariensis} Mantadia NP, East Madagascar
Leopard (Panthera pardus) male climbing on fallen tree, , Nakuru national park, Kenya
Beluga whale (Delphinapterus leucas) play biting camera, Arctic circle Dive Center, White Sea, Karelia, northern Russia. Captive
Silhouette of Red-fronted brown lemur (Eulemur rufus) leaping across gap in trees by moonlight, cathemeral behaviour, Ranomafana National Park, south east Madagascar, November
Head portrait of Satiny Parrot snake (Leptophis depressirostris) swallowing frog, Hungary, Europe
Gannet (Morus bassanus) courtship, Saltee Islands, County Wexford, Ireland, June 2009
Female African lion (Panthera leo) jumping in to the Khwai river, Okavango Delta, Moremi reserve, Botswana, sequence 1/5
Brown bear {Ursus arctos} enjoys hot steam from a geyser in Valley of the Geysers, Kronotsky Zapovednik, Kamchatka, Far East Russia
Meerkat family group at burrow {Suricata suricatta} Tswalu Kalahari Reserve South Africa
Impala (Aepyceros melampus) male, Masai-Mara Game Reserve, Kenya
Cheetah (Acinonyx jubatus) cubs aged 6 / 7 weeks old playing, Masai-Mara Game Reserve, KenyaCheetah (Acinonyx jubatus) cubs aged 6/7 weeks old playing, Masai-Mara Game Reserve, Kenya. Vulnerable species
Wildebeest (Connochaetes taurinus) migration, herd crossing the Mara river, Masai-Mara Game Reserve, Kenya
Agama (Agama agama) male, Tsavo East National Park, Kenya
Herd of Southern giraffes (Giraffa camelopardalis giraffa) and Burchells zebras (Equus quagga) aerial view, Okavango delta, Bostwana
Bottlenose dolphin (Tursiops truncatus) playing in the waves of an oil tanker, Port Aransas, Texas, USA
African Buffalo (Syncerus caffer) males fighting, Nakuru National park, Kenya
Bridled parrotfish (Scarus frenatus) clownish grin reveals its power tools: grinding teeth used to scrape algae from rock, Maldives, Indian Ocean
Beluga whale (Delphinapterus leucas) swimming under ice and exhaling air, with scuba diver in background, Arctic circle Dive Center, White Sea, Karelia, northern Russia, captive
Atlantic Puffin (Fratercula arctica) resting on a snow bank, Isle of Hornoya, Varanger, Norway, April. Bookplate from Danny Greens The Long Journey North
Fulmer (Fulmras glacialis) in flight near blue glacier, Svalbard, July. Bookplate from Danny Greens The Long Journey North
White tailed Sea Eagle (Haliaeetus albicilla) in flight, Norway, July. Bookplate from Danny Greens The Long Journey North
Tawny Owl (Strix aluco) adult female hiding behind tree trunk, trained bird, Somerset, UK, January
Split image of Blue shark (Prionace glauca) close to the surface, Pico Island, Azores, Portugal, Atlantic Ocean, July
Shoal of Longfin bannerfish (Heniochus acuminatus) Maldives, Indian Ocean
Shoaling glassfish (Parapriacanthus sp. ) shoal, Maldives, Indian OceanShoaling glassfish (Parapriacanthus sp.) shoal, Maldives, Indian Ocean
Lammergeier (Gypaetus barbatus) close up portrait, Spain, November
Capercaillie (Tetrao urogallus) in flight, Kuusamo, Finland, January
Three Dalmatian Pelicans (Pelecanus crispus) portrait on lake. Lake Kerkini, Greece, March 2012
Eurasian Lynx (Lynx lynx) adult sitting on boulder covered in deep snow in beech (Fagus sylvatica forest, captive in enclosure of the Bavarian Forest National Park, Germany, February
Harvest mouse (Micromys minutus) on barley cereal, Yorkshire, UK Captive
Northern gannets (Morus / Sula bassanus) portrait of individual with a courting pair in foreground, UK, Bass Rock, Scotland
Andean condor (Vultur gryphus) in flight over mountain peaks, Torres Del Paine National park, Chile
Blue Whales (Balaenoptera musculus) throat inflated, filter feeding at sea surface. California, USA. Sequence 1 of 2
Humpback Whale (Megaptera novaeangliae) calf, Tonga, South Pacific, September
Humpback Whale (Megaptera novaeangliae) mother and calf. Tonga, South Pacific, September
Portrait of a Yellow seahorse (Hippocampus guttulatus), Capo Galera, Alghero, Sardinia, Italy, Mediterranean Sea
Bohemian waxwings (Bombycilla garrulus) feeding on berries, Uto Finland October
Ruff (Philomachus pugnax) two males displaying in territorial dispute, Liminka Finland June
Golden Eagle (Aquila chrysaetos) portrait in snow, Utajarvi, Finland February
Arctic Tern (Sterna paradisaea) in flight, silhouetted against the moon at dusk, Uto Finland July
Polar Bear (Ursus maritimus) head portrait with blue tongue sticking out, Svalbard, Norway
White rhinoceros (Ceratotherium simum) mother and calf browsing. Entabeni Safari Conservancy, Limpopo region, Waterberg, South Africa, October
Two Red Foxes (Vulpes vulpes) fighting. The Netherlands. August
Male Marsh pride lion (Panthera leo) on a moonless night, Masai Mara, Kenya. Taken with infra red camera, September
Snow Geese (Chen caerulescens) in flight, silhouetted against colourful dusk sky. Bosque del Apache, New Mexico, USA, November
Snow Goose (Chen caerulescens), coming in to land. Bosque del Apache, New Mexico, USA, November