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Marine Iguana (Amblyrhynchus cristatus) on rocks with wave splash. Galapagos, Ecuador
Red Squirrel (Sciurus vulgaris) adult reaching up. Cairngorms National Park, Scotland, UK, March
Heermanns Gulls (Larus heermanni) in flight. Ensenada, Baja California Peninsula, Mexico, December
American Flamingo (Phoenicopterus ruber), Ria Lagartos Biosphere Reserve, Yucatan Peninsula, Mexico, August
Polar bear (Ursus maritimus) adult walking over newly formed pack ice during fall freeze up, 1002 area of the Arctic National Wildlife Refuge, North Slope, Alaska, USA
Polar bear (Ursus maritimus) sow and cub sliding on their backs over the pack ice, much like ice skating, 1002 coastal plain of the Arctic National Wildlife Refuge, Alaska, USA
Dall sheep (Ovis dalli) solitary animal at sunset with a storm moving across the sky, Denali National Park, interior Alaska, USA
Salmon landing on head of Grizzly bear (Ursus arctos horribilis) as it is leaping up rapids, Katmai National Park, Alaska, USA
Long Horn cattle in meadow, Norfolk, UK July
A school of lesser devil rays (Mobula hypostoma) flying through sunbeams as they feed. Isla Mujeres, Quintana Roo, Yucatan Peninsula, Mexico. Caribbean Sea
Whitetip reef sharks (Triaenodon obesus) pack silhouetted following scent trail in water column, Cocos Island, Costa Rica, Pacific Ocean
Emperor angelfish (Pomacanthus imperator) with soft corals and sweepers. Egypt, Red Sea
Spectacled caiman (Caiman yacare) walking on a sandy shore at the edge of a river at night, Pantanal, Brazil
Little egret (Egretta garzetta) attempting to catch fish, Camargue, France, July
White terns (Gygis alba) flock in flight overhead, Christmas Island / Kiritimati, Pacific Ocean, July
Killer WHale / Orca (Orcinus orca) bull and cow in Resurrection Bay, Kenai Fjords National Park, outside Seward, Alaska, May
Great Egrets (Ardea alba) territorial fight above nest colony, Everglades National Park, Florida, USA, February
Whooper swan (Cygnus cygnus) in snowfall. Martin Mere Wetlands Trust, Lancashire, UK, January. Animal Portraits category, British Wildlife Photographer Awards (BWPA) competition 2012
White bellied woodstar (Chaetocercus mulsant) three hovering around red flowers, Guango private reserve, Papallacta Valley, Andean Cloud Forest, East slope, Ecuador
Purple gallinule (Porphyrio martinicus) moving across waterlily covered surface, Everglades National Park, Florida, USA, March
Whiskered auklet (Aethia pygmaea), Iony Island / Jonas Island, Sea of Okhotsk, Far East Russia, July
Flock of Greater flamingo (Phoenicopterus ruber) in flight, Camargue, Southern France, November 2006
Robin (Erithacus rubecula) perched on rosehip branch during snowfall. UK, December
Black faced Impala (Aepyceros melamis petersi) female jumping, Etosha National Park, Namibia
Cowboys rounding up a band of Criollo pure pedigree mares and foals, Estancia Ita Maria, Misiones, Paraguay, January 2012
A band of semi-feral Quarter mares and foals running across a creek, Estancia Don Amerigo, Chaco, Paraguay, January 2012
Malachite kingfisher (Alcedo cristata) perched on rush, Intaka Island, Cape Town, South Africa, February
Common peafowl (Pavo cristatus) perched on trees at dawn, Western Ghats, Southern India
Close-up of Brown bear (Ursus arctos) in lake, Kamchatka, Far east Russia, August
Asiatic lionesses and cubs (Panthera leo persica) drinking from pool, Gir Forest NP, Gujarat, India. Water is scarce in the Gir
Mute swans (Cygnus olor) newly hatched cygnets on pond, Yetholm Loch Scottish Wildlife Trust Reserve, Roxburghshire, Scotland, May
Highland bull (Bos taurus) face close up in freezing fog, Berwickshire Scotland, December
Goldfinches (Carduelis carduelis) squabbling near seed feeder, Berwickshire, Scotland, April
Brown trout (Salmo trutta) resting on river bed near the Aigas Field Studies Centre, Inverness-shire, Scotland, July
Clouded leopard (Neofelis nebulosa) Guwahati, Assam, India, captive
Mistle Thrush (Turdus viscivorus) in snow. Hertfordshire, England, UK, February
Southern plains grey langur ( Semnopithecus entellus / Presbytis entellus) close up face profile, Sawai Modhopur, Rajasthan, India
Boxer looking over closed door, standing on hind legs
Spotted scorpionfish (Scorpaena plumieri) lying motionless and camouflaged in soft corals. West Palm Beach, Gulf Stream, West Atlantic Ocean, Florida, USA
Banded jawfish (Opistognathus macrognathus) male incubating eggs in mouth, which are at a late stage of development and will soon be ready for release
Yellow goatfish (Mulloidichthys martinicus) swimming above the reef, West End, Grand Cayman, Cayman Islands, British West Indies, Caribbean Sea
Spotted owl (Strix nebulosa) Gifford-Pinchot National Forest, Washington USA
Grey Heron (Ardea cinerea) taking a fish from water. Kiskunsagi National Park, Hungary, May
Two large white piglets (Sus scrofa domestica) at 3 weeks in sty, UK
Ruffs displaying and fighting (Philomachus pugnax) Agapa River, Taimyr Peninsula, Siberia, Russia
Giraffe (Giraffa camelopardalis) three standing together, silhouetted at dusk, Okavango Delta, Botswana
Freerange Gloucester old spot domestic pig (Sus scrofa domestica) portrait with ears covering eyes, UK
Cotswolds Lion rare breed sheep (Ovis aries) and the village of Naunton at sunset, Gloucestershire, UK, June 2001
