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Caribbean reef shark (Carcharhinus perezi) over sand ripples, Walkers Cay, Bahamas. Tropical West Atlantic Ocean. Taken in natural light
Red deer (Cervus elaphus) stag amongst winter vegetation, Salburua Park, Alava, Spain, November
African lion (Panthero leo) male giving flehmen grimace, Tanzania, East Africa
Little owl (Athene noctua) perched on post at dusk, Carmarthenshire, Wales, UK, Captive
Feral domestic horses (Equus caballus) wild herd stallion (grey) fighting a bachelor chestnut stallion, Blue Swallow Reserve, near Kaapsehoop, Mpumalanga province / Eastern Transvaal, South Africa
Polar bears (Ursus maritimus) female with cubs aged two years travelling along a barrier island during autumn freeze up, Bernard Spit, 1002 area of the Arctic National Wildlife Refuge, Alaska
Harlequin duck (Histrionicus histrionicus) portrait of male stood motionless on exposed rock, in river, Iceland, May
Two African lion cubs (Panthera leo) playing on a road in the Kalahari Desert, Botswana
Old male African lion (Panthera leo) portrait, Sabi Sand Game Reserve, South Africa, June
Catesbys snail-eating snake (Dipsas catesbyi) on Heliconia plant, Pacaya Samiria National Reserve, Amazon Basin, Loreto, Peru
Laysan albatross (Phoebastria immutabilis) head portrait calling mate, Guadalupe Island Biosphere Reserve, off the coast of Baja California, Mexico, March
Northern elephant seal (Mirounga angustirostris) close-up of weaner / juvenile sleeping, Guadalupe Island Biosphere Reserve, off the coast of Baja California, Mexico, April
African elephant (Loxodonta africana) at waterhole with Giraffe in the background, Etosha National Park, Namibia, August
European hare (Lepus europaeus) mating pair boxing in field, Slovakia
Panther Chameleon {Furcifer pardalis} walking along branch on coast, Masoala Peninsula National Park, north east Madagascar
Herd of Wildebeest (Connochaetes taurinus) silhouetted at dusk, Serengeti NP, Tanzania, East Africa, January
An adult female California sea lion (Zalophus californianus) head portrait, in a rocky underwater cave. La Paz, Baja California Mexico. Sea of Cortez, East Pacific Ocean. October
California sea lion pup (Zalophus californianus) portrait in a rocky underwater cave. La Paz, Baja California Mexico. Sea of Cortez, East Pacific Ocean. October
Atlantic grey seal (Halichoerus grypus) swimming beneath the surface, Lundy Island, Devon, England, UK. July 2010. Highly commended, Coast and Marine category
Group of Manta rays (Manta birostris) feeding together on plankton in a shallow lagoon. Hanifaru Lagoon, Baa Atoll, Maldives. Indian Ocean
Black browed albatross (Thalassarche melanophrys) flying in 60 knot wind at Cape Horn, Chile
A group of Florida manatees (Trichechus manatus latirostrus) sleeping in the afternoon at Three Sisters Spring. Crystal River, Florida, USA
Lemon shark (Negaprion brevirostris) in shallow water with reflection at the surface. Little Bahama Bank, Bahamas
Caribbean reef shark (Carcharhinus perezi) swims over a coral reef. Grand Bahama Island, Bahamas. Tropical West Atlantic Ocean
Bottlenose dolphins (Tursiops truncatus) in shallow water over a sand bank. Sandy Ridge, Little Bahama Bank, Bahamas. Tropical West Atlantic
A pod of Atlantic spotted dolphins (Stenella frontalis) riding on the bow wave of a boat. Sandy Ridge, Little Bahama Bank. Bahamas
Polar bear (Ursus maritimus) footprints in the snow along a barrier island during autumn freeze up, Bernard Spit, 1002 area of the Arctic National Wildlife Refuge, Alaska. October 2009
Polar bear (Ursus maritimus) head close-up portrait of an adult male, with snowflakes on fur, Barter Island, 1002 area of the Arctic National Wildlife Refuge, Alaska
Common / Eurasian cranes (Grus grus) small flock of juveniles taking flight over the frozen, snow covered pastureland on a foggy winter morning
Red Kite (Milvus milvus) diving towards prey whilst in flight, Wales, UK, March
Dipper (Cinclus cinclus) landing on exposed stone, with insect prey for chicks in beak, Brecon Beacons National Park, Wales, UK
Crested Newt (Triturus cristatus carnifex) underwater view of female swimming on surface, Traversari lake, Campigna National Park, Emilia Romagna and Tuscany regions, Italy
Sri Lankan Leopard (Panthera pardus kotiya) soft focus running, captive
As soon as it entered the water the young Olive ridley sea turtle (Lepidochelys olivacea) struggles against the swell to swim away from the coast
These hatchlings of an Olive ridley sea turtle (Lepidochelys olivacea) should be buried and protected under 40 cm of warm sand, but a domestic dog dug them out at Ostional beach in Costa Rica
Mountain hare (Lepus timidus) running across wind swept snow field, near Carrbridge, Cairngorms National Park, Scotland, January 2010
Kromfohrlander running / racing in field towards camera
Small Munsterlander head portrait
Dalmatian, head portrait, male aged 4 years
Alaskan Malamute portrait of blue coated adult
Golden damselfish (Amblyglyphidodon aureus) swimming past coral reef wall with colourful soft corals. Bali, Indonesia
Fieldfare (Turdus pilaris) feeding on Rowan berries (Sorbus hupehensis) in winter, Perthshire, Scotland. WINNER: Young Wildlife Photographer of the Year 2010
Adult male Bornean orangutan (Pongo pygmaeus) in rainforest canopy, Gunung Palung National Park, Borneo, West Kalimantan, Indonesia
Flock of Cattle Egrets (Bubulcus ibis) in roosting trees at dusk, Imba Ndutu, Serengeti NP, Tanzania, East Africa
Arctic tern (Sterna paradisaea) in flight, vocalising, aggresive behaviour, Farne Islands, Northumberland, UK, June
Arctic tern (Sterna paradisaea) in flight, calling, Farne Islands, Northumberland, UK, June
Large group of Californian sealions (Zalophus californianus) swimming underwater, Anacapa Island, California, Pacific Ocean
Cheetah (Acinonyx jubatus) running at full speed chasing Springbok prey, Kalahari Desert, Southern Africa
