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Whooper swans (Cygnus cygnus) two in flight, during snowfall, Lake Kussharo, Japan, February
Yellow-plumed parakeets (Leptosittaca branickii) at nest cavity in wax palm stump, Tapichalaca Biological Reserve, Ecuador
Barn owl on post in flooded Somerset Levels. England, UK, Europe
Bateleur eagle (Terathopius ecaudatus) head portrait, Captive, occurs in Africa
Red squirrel (Sciurus vulgaris) having a drink, Black Isle, Scotland, UK. October
Long-eared owl (Asio otus) in a large tree, Cheshire, UK, March
Backlit photo of a Robust ghost pipefish (Solenostomus cyanopterus) reveals a small fish that it has just eaten in its stomach. Lembeh Strait, North Sulawesi, Indonesia. Molluca Sea
Wren (Troglodytes troglodytes) on frost covered fern, Dunham Massey, Cheshire, UK. Winner of the Habitat Category of the British Wildlife Photographer of the Year Awards (BWPA) 2017
Brandts cormorant (Phalacrocorax penicillatus) bursts through a school of Pacific chub mackerel (Scomber japonicus) while hunting beneath an oil rig
Corys shearwaters (Calonectris diomedea) diving among a mass of shoaling fish to feed, along with Atlantic spotted dolphins (Stenella frontalis), Formigas Islet dive site, Azores, Portugal
Adelie penguin (Pygoscelis adeliae) diving near ice flow, Antarctica. Small reproduction only
Desmarests hutia (Capromys pilorides doceleguas), Jardines de la Reina / Gardens of the Queen, Caribbean Sea, Ciego de Avila, Cuba, January
Himalayan monal (Lophophorus impejanus) male perched on branch, Uttarakhand, India
Golden eagle (Aquila chrysaetus) Kuusamo, Finland, January
Coal tit (Periparus ater) on a branch with lichen, Vendee, France, December
Portrait of a Green sea turtle (Chelonia mydas) in morning light. Sipadan Island, Sabah, Borneo, Malaysia. Celebes Sea
Two Sulawesi black macaques / Celebes crested macaques (Macaca nigra) juveniles, in woodland, portrait, Tangkoko National Park, northern Sulawesi, Indonesia. Critically endangered
Golden plover (Pluvialis apricaria) chick camouflaged in ground nest, Norway June
Three Red-crowned cranes (Grus japonensis) in flight. Hokkaido, Japan. March
Pelagic thresher shark (Alopias pelagicus) swimming in open ocean, Malapascua Island, Philippines, Pacific Ocean
Common loon (Gavia immer), in breeding plumage, running across water to take flight, Michigan, USA. June
Bison (Bison bison) female with calf walking through snow in front of frost-covered forest, Yellowstone National Park, USA
Caspian Tern (Sterna caspia) in flight, diving for prey. Lake Ellesmere, Canterbury, South Island, New Zealand
Two Red-footed booby (Sula sula) perched on branches, Genovesa Island, Galapagos Islands, Ecuador
White tailed eagle (Haliaeetus albicilla) calling in flight, Isle of Mull, Argyll Scotland, UK. October
Spotted eagle rays (Aetobatus narinari) swimming, Hawaii, USA
Common buzzard (Buteo buteo) in flight with hoar frost. Northamptonshire, UK, January
Barbary macaque (Macaca sylvanus), two adults with baby, overlooking town and ships in sea at sunset. Gibraltar Nature Reserve, Gibraltar. August 2018
Red-and-Green Macaws (Ara chloropterus), feeding at the wall of a riverside clay lick. Blanquillo Clay Lick, Manu Biosphere Reserve, Peru. November
Bottlenose dolphins (Tursiops truncatus) at the surface at sunrise. Kwazulu-Natal, South Africa
Golden eagle (Aquila chrysaetus), Kuusamo, Finland, January
Common loon (Gavia immer) nesting at edge of lake and calling, Acadia National Park, Maine, USA, August
Brown bear (Ursus arctos arctos) female and two cubs, aged 1 year, emerging from den in winter, Bavarian Forest, Germany, captive, January
1468136 - Mandrill male (Mandrillus sphinx) abstract of brightly coloured ischial1468136 - - Mandrill male (Mandrillus sphinx) abstract of brightly coloured ischial callosity on buttocks, Lekedi National park, Gabon
Cheetah (Acinonyx jubatus) staring back over its shoulder, black and white. Save Valley Conservancy, Zimbabwe, August
Aerial view of Reindeer (Rangifer tarandus) fording water whilst being herded by Nenet herders. Nenets Autonomous Okrug, Arctic, Russia, July 2017
RF- Carrion crow (Corvus corone) flying from a snow covered pine tree in a winter landscape. Valsavarenche, Gran Paradiso NP, Italy, March
Clarks Grebe (Aechmophorus clarkii) with yawning chick riding on back, Bear River Refuge, Utah, July
Ptarmigan (Lagopus muta) male on snowy ridge on Cairngorm mountain, Cairngorms National Park, Scotland, UK, March
White-throated dipper (Cinclus cinclus) winter with river flooding, Kuusa, Laukaa, Keski-Suomi, Lansi- ja Sisa-Suomi / Central and Western Finland, Finland. January
Pheasant (Phasianus colchicus), plumage detail. UK, March
Violet sabrewing hummingbird (Campylopterus hemileucurus) male hummingbirds territorial fight at dawn, Turrialba, Costa Rica
Great crested grebe (Podiceps cristatus) pair performing courtship displaying at dawn, backlit and surrounded by mist, Cheshire, UK, April
Gentoo penguins (Pygoscelis papua) group walking along snow, Cuverville Island. Antarctic Peninsula, Antarctica
Coastal brown bear (Ursus arctos) and cub, Lake Clarke National Park, Alaska, September
Grey Seal (Halichoerus grypus) at rest on submerged rock, head and one flipper above water. Shetland Islands, Scotland, UK. July
Raven (Corvus corax) calling in the snow, Kemijarvi, Finland, February
Buzzard (Buteo buteo) in flight, portrait, Marlborough Downs, Wiltshire, UK. December