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Grayling (Thymallus) migrating to spawning in the Temnik River, Lake Baikal, Baikalsky Reserve, Siberia, Russia. May
Murmuration of Common starlings (Sturnus vulgaris) with Peregrine falcon (Falco peregrinus) hunting them, The Netherlands, November
Fin whale (Balaenoptera physalus) near surface. Pico Island, Azores, Portugal. May
Orcas / killer whales (Orcinus orca) swimming in open water, Three Kings Islands, New Zealand. Pacific Ocean
Bearded pig (Sus barbatus) close up of snout, Bako National Park, Sarawak, Borneo
Whooper swan (Cygnus cygnus) in landscape of Landmannalaugar, Iceland, June
A portrait of a male Ballan wrasse (Labrus bergylta), showing his bright mating colours. Plymouth, Devon, England, Great Britain. Wreck of the Glen Strathallan, Plymouth Sound, English Channel
Bornean Orangutan (Pongo pygmaeus) female face portrait, Tanjung Puting reserve, Camp Leakey, Central Kalimantan, Borneo
Male Black rhinoceros (Diceros bicornis) running, Nakuru National Park, Kenya
Mountain gorilla (Gorilla gorilla beringei) silverback Gihishamwotsi displaying, non group dominant, Sabyinyo Group, Volcanoes National Park, Virunga Mountains, Rwanda
Leopard (Panthera pardus) female fighting off male after he tries to mate with her. Londolozi Private Game Reserve, Sabi Sand Game Reserve, South Africa
Whooper swans (Cygnus cygnus) three coming into land, Hokkaido, Japan, February
Grey whale (Eschrichtius robustus) at water surface with mouth open showing baleen plates, San Ignacio Lagoon, Baja California, Mexico
Harvest mouse (Micromys minutus) climbing among Cow Parsley, Hertfordshire, England, UK, May, Controlled conditions
Great Bustard (Otis tarda) males displaying, Salisbury Plain, Wiltshire, UK, April
Swallow-tailed gull (Creagrus furcatus) Plazas Island, Galapagos
RF - Snow leopard (Panthera uncia) female, portrait, captive
Snow leopard (Uncia uncia) Altai Mountains, Mongolia. March
Robin (Erithacus rubecula) at RSPB Leighton Moss Bird Reserve, Silverdale, Lancashire, England, UK. February 2016
Landscape of Zanskar River, a tributary of the Indus, borders part of Hemis national park, habitat of the snow leopard (Panthera uncia), Hemis National Park, Ladakh, India
Yellow bellied toad (Bombina variegata) showing underside, Kirchheimbolanden, Germany. Meetyourneighbours.net project
Yellow Wattlebird (Anthochaera paradoxa) Tasmania, Australia, endemic to Tasmania
Caribou / Reindeer bull (Rangifer tarandus) close-up grazing, Denali National Park, Alaska, USA, September
Moose (Alces alces) bull swimming in water, Baxter State Park, Maine, USA
American bison (Bison bison) Yellowstone National Park, Montana, USA
Grizzly bear (Ursus arctos) catching a fish, Brooks Falls in Katmai National Park, Alaska, USA, July
Red deer (Cervus elaphus) stag silhouetted at sunset, Scotland, UK, February
Black-throated Diver (Gavia arctica) Rokua, Finland, May
European hare (Lepus europaeus), Wirral, England, UK, May
Jaguar (Panthera onca) male walking, Pantanal, Brazil
Scarlet macaws (Ara macao) La Selva, Costa Rica
Silhouette of Common peafowl (Pavo cristatus) perched in a tree at dawn. Keoladeo Ghana National Park, Bharatpur, Rajasthan, India
Arctic Fox (Alopex / Vulpes lagopus) standing on ridge, during moult from grey summer fur to winter white. Dovrefjell National Park, Norway, September
Arctic Fox (Alopex / Vulpes lagopus) yawning, during moult from grey summer fur to winter white. Dovrefjell National Park, Norway, September
Japanese cranes (Grus japonensis) displaying in snow, Hokkiado, Japan, February
Willow Grouse (Lagopus lagopus) male perched on snow, Finland, March. Bookplate from Danny Greens The Long Journey North
Lions (Panthera leo) - two brothers patrolling territorial boundary, affectionate behaviour, border of Serengeti / Ngorongoro Conservation Area (NCA) near Ndutu, Tanzania
Stellers sea-eagle (Haliaeetus pelagicus) landing on pack ice, Hokkaido, Japan, February
Pine marten (Martes martes) reflected in water, Ardnamurchan Peninsula, west coast of Scotland, UK. Highly commended in the Mammals category of the British Wildlife Photography Awards (BWPA)
Weasel (Mustela nivalis) looking out of hole on woodland floor with snowdrops, Sheffield, England, UK
Young urban Red fox (Vulpes vulpes). Bristol, UK. August
RF- Goldfinch (Carduelis carduelis) reflected in pool, Worcestershire. November
Barn owl (Tyto alba) in flight, captive, Cumbria, UK
A portrait of a Mediterranean parrotfish (Sparisoma cretense) above an algae covered rock. El Hierro, Canary Islands, Spain. East Atlantic Ocean
School of Silversides (Atherinomorus lacunosus) mass below a jetty, creating a false ceiling of fish. Berenice Jetty, Aqaba, Jordan. Gulf of Aqaba, Jordan
Dalmatian Pelican (Pelecanus crispus) portrait, close-up of eyes, Lake Kerkini, Greece. February
Siberian tiger (Panthera tigris altaica) in snow, captive
Siberian musk deer (Moschus moschiferus) male in snow, Irkutsk, Russia. January
