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Eastern chipmunk (Tamias striatus) gathering chestnuts, Chestnut Hill, Philadelphia, Pennsylvania, USA, September
Stellers sea-eagle (Haliaeetus pelagicus) in flight, Hokkaido, Japan, February
Galapagos penguin (Spheniscus mendiculus) underwater, Bartholome Island, Galapagos Islands, East Pacific Ocean
Angular roughshark (Oxynotus centrina) a deepsea species living at 80-300m depth, caught by fishermen and released, Costa Brava, Catalunya, Spain, Mediterranean Sea
Plains zebra (Equus quagga) grooming foal, Kruger National Park, South Africa, July
Giant panda cub (Ailuropoda melanoleuca) portrait Yuan Meng, first giant panda ever born in France, age 10 months. Captive at Beauval Zoo, Saint Aignan sur Cher, France
Man-of-war fish (Nomeus gronovi) living within deadly tentacles of Portuguese man o war (Physalia physalis), to whose toxin it is ten times more resistant than other fish, at sea surface. Dominica
Australian sea (Neophoca cinerea) juvenile playing in the sand, Carnac Island, Western Australia
Smooth hammerhead shark (Sphyrna zygaena). Cabo San Lucas, Baja California Sur, Mexico
Red deer (Cervus elaphus) backlit at dawn with visible breath. UK. October
Flock of ducks silhouetted against sunset flying over field in winter in the Uitkerkse Polder nature reserve near Blankenberge, West Flanders, Belgium. February 2020
Snow leopard (Panthera uncia) female in snow, Hemis National Park, Ladakh, India
Great Spotted Woodpecker in snow (Dendrocopos major), Scotland, February
Brown bear (Ursus arctos) male, portrait. Martinselkonen, Kainuu, Finland. June
Bamboo lemur (Hapalmur griseus) in tree. Andasibe-Mantadia National Park, Madagascar
Short-beaked echidna (Tachyglossus aculeatus). Francois Peron National Park, Shark Bay, Western Australia. October
Japanese crane (Grus japonensis) displaying, wings in heart shape, Hokkiado, Japan, February
Kaleidoscopic image of Clarks anemonefish (Amphiprion clarkii), Ambon, Indonesia
Sable (Martes zibellina) in tree, Irkutsk, Siberia, Russia. November
Roughtail stingray (Dasyatis centura) on sea floor, Tenerife, Canary Islands
Red fox (Vulpes vulpes) in winter snow, Jura, Switzerland
Blue-footed booby (Sula nebouxii) landing, South coast, Santa Cruz Island, Galapagos
Great frigatebird (Fregata minor) in tree with pair of Nazca boobies (Sula granti) showing mating behaviour, Genovesa Island, Galapagos
Red-billed tropicbird (Phaethon aethereus) in flight, Plazas Island, Galapagos
Eastern tiger salamander (Ambystoma tigrinum) North Florida, USA. December
Great spotted woodpecker (Dendrocopos major), male feeding juvenile in nest, Finland, June
Eastern lowland gorilla (Gorilla beringei graueri) silverback named Chimanuka climbing a tree, Kahuzi-Biega National Park, South Kivu Province, Democratic Republic of Congo (DRC)
Canadian timber wolf / Northwestern wolf (Canis lupus occidentalis) pack moving through snow, Wood Buffalo National Park, Alberta, Canada. Taken on location for BBC Frozen Planet, March 2009
House Martin (Delichon urbica) Spain, April
Great Grey Owl (Strix nebulosa) hunting over snow, Kuhmo Finland, March
Humpback whale (Megaptera novaeangliae) male calf relaxing with his mother and an escort whale in the background. Vava'u, Tonga, South Pacific
Arctic tern (Sterna paradisaea) hovering above the ocean, Svalbard, Norway June
Australian sea lion (Neophoca cinerea) playful juvenile swimming upside-down, Carnac Island, Western Australia
American Pika (Ochotona princeps) leaping from one alpine rock to another as it heads to its home carrying food for the winter months. Rocky Mountain National Park, Colorado, USA. August
Kestrel (Falco tinnunculus) male hovering overhead, Cornwall, UK, April
Arctic fox (Vulpes lagopus) portrait of juvenile, winter pelage. Dovrefjell National Park, Norway. September
Spotted-tailed quoll (Dasyurus maculatus) scent marking in Monga National Park, New South Wales, Australia. Remote camera, triggered by movement
Kakapo (Strigops habroptilus) close up showing sensory facial feathers, Codfish Island / Whenua Hou, Southland, New Zealand, February. Critically endangered
Common eider duck (Somateria mollissima) female defending her nest from arctic fox (Vulpes lagopus). Longyearbyen, Spitsbergen. June
Humpback whale (Megaptera novaeangliae) just under surface, off Shetland, Scotland, UK. First ever underwater images of humpbacks in British waters
Black howler (Alouatta caraya) captive, occurs in South America
Young Platypus (Ornithorhynchus anatinus) is released onto a log in McMahons Creek, Yarra Ranges, Victoria, Australia. Photographed under controlled conditions. September
Arctic fox (Vulpes lagopus), juvenile running through snow, winter pelage. Dovrefjell National Park, Norway. September
Arctic fox (Vulpes lagopus) juvenile sniffing ground, winter pelage. Dovrefjell National Park, Norway. September
Red fox (Vulpes vulpes) at night in snow, camera trap image, Jura Mountains, Switzerland, August. Commended in the Mammals category of the Asferico Competition 2016
Australasian crested grebe pair (Podiceps cristatus australis) in courtship display. Ashburton Lakes, Canterbury, New Zealand. August
Great crested grebe (Podiceps cristatus) performing their weed dance during courtship. Den Oever, The Netherlands. July
Marine iguana (Amblyrhynchus cristatus) swimming underwater, Fernandina Island, Galapagos