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Santa Cruz Galapagos tortoise (Chelonoidis nigra porteri) feeding on grass, Santa Cruz Highlands, Galapagos
African lion (Panthera leo) portrait, Masai Mara Game Reserve, Kenya
Emperor penguin (Aptenodytes forsteri) viewed through hole in iceberg at Snow Hill Island rookery, Antarctica. November
Eurasian flamingos (Phoenicopterus roseus) lined up on Pulicat Lake, Tamil Nadu, India, January 2013
Gannets (Morus bassanus) diving to feed on discarded fish, Shetland, Scotland, UK, April
Female Harpy Eagle (Harpia harpyja) in flight. Returning to its nest. Pousada Currupira d Araras, south west Brazil
Puffin (Fratercula arctica) standing on flowering Sea thrift (Armeria maritima). Portrait. Fair Island, Shetland Islands, Scotland, July
RF - Southern stingray (Dasyatis americana) swimming over sand in shallow water at dawn, Cayman Islands, Caribbean Sea
Sea otter (Enhydra Lutris) with forelegs raised, Sitka Sound, Alaska, USA, August
Great hammerhead shark (Sphyrna mokarran) mouth wide open, feeding in shallow water, surrounded by fish. South Bimini Island, Bahamas
Red deer (Cervus elaphus) stag in upland landscape. Lochcarron, Highlands, Scotland, UK
Arctic fox (Vulpes lagopus), juvenile looking at camera, portrait, winter pelage. Dovrefjell National Park, Norway. September
Head portrait of black Merens stallion with long mane running in pasture. Northern France, Europe. February
Common kestrel (Falco tinnunculus) perched on a branch, Valencia, Spain, February
Raccoon (Procyon lotor) peering out from behind tree trunk, Baxter State Park, Maine, USA
Grey Wolf (Canis lupus), in forest, captive, USA
Close up portrait of the face of a Nurse shark (Ginglymostoma cirratum) resting on the sand in shallow water. Its barbels are clearly visible on its top lip. South Bimini, Bahamas
Japanese macaque (Macaca fuscata) sleeping at hot spring in Jigokudani, Yaenkoen, Nagano, Japan, February
Cuckoo (Cuculus canorus) perched on a branch. Thursley Common, Surrey, UK, May
Polar bear cubs (Ursus maritimus) triplets age 2-3 months next to their mother. Wapusk National Park, Manitoba, Canada. February
Portrait of a Mandarin duck (Aix sponsa) male animal and female. UK. Introduced species
Red sea fan (Melithaea sp.) is surrounded by Glassfish ( Apogon sp.) on a coral reef. Daram Islands, Misool, Raja Ampat, West Papua, Indonesia. Ceram Sea. Tropical West Pacific Ocean
Peregrine falcon (Falco peregrinus) in flight over cotton grass flowers, Vaala, Finland, June
Siberian flying squirrel (Pteromys volans orii) flying directly over photographer. Hokkaido, Japan. February
Indo-Pacific bottlenose dolphin (Tursiops aduncus) with penis extended. Ogasawara / Bonin Islands, Japan
Bald eagle (Haliaeetus leucocephalus) pair flying with claws linked during courtship flight, Homer, Alaska, USA, March
RF- Jaguar (Panthera onca) head portrait, captive
RF- White horse of the Camargue, running from sea. Camargue, Southern France
Portrait of a Golden snub-nosed monkey (Rhinopithecus roxellana) screaming and showing its teeth, Foping Nature Reserve, Shaanxi, China. Endangered species
Indian peafowl (Pavo cristatus) peacock displaying feathers, captive, occurs in South Asia
RF - Arctic Fox (Vulpes lagopus) portrait in winter coat, Svalbard, Norway, April
Orcas / killer whales (Orcinus orca) swimming in open water, Three Kings Islands, New Zealand. Pacific Ocean
Giant Manta Ray (Manta birostris), San Benedicto Island, Revillagigedo Archipelago Biosphere Reserve (Socorro Islands), Pacific Ocean, Western Mexico. Vulnerable species
Tompot blenny (Parablennius gattorugine) portrait, Swanage Pier, Swanage, Dorset, UK, English Channel, August
Pod of Indian Ocean bottlenose dolphin (Tursiops adunctus) swim over a coral reef. Sha ab El Erg, Hurghada, Egypt. Red Sea
RF - Keel-billed toucan (Ramphastos sulfuratus) perched on mossy branch. Boca Tapada, Costa Rica. (This image may be licensed either as rights managed or royalty free.)
Male Fossa (Cryptoprocta ferox) climbing down tree trunk from forest canopy. Mid-altitude rainforest, Andasibe-Mantadia National Park, eastern Madagascar. Endangered
Red-footed booby (Sula sula) in flight with wings outstretched. Genovesa Island, Galapagos
Giant anteater (Myrmecophaga tridactyla) Serra de Canastra National Park, Brazil
Barn Swallow (Hirundo rustica) in flight. Portugal, Europe
Shoebill stork (Balaeniceps rex) in the swamps of Mabamba, Lake Victoria, Uganda
Killer whale (Orcinus orca) pod hunting together in herring baitball (Clupea harengus), Andenes, Andoya island, North Atlantic Ocean, Norway, April
A portrait of a male Ballan wrasse (Labrus bergylta), showing his bright mating colours. Plymouth, Devon, England, Great Britain. Wreck of the Glen Strathallan, Plymouth Sound, English Channel
Head portrait of grey Arabian stallion running in desert dunes near Dubai, United Arab Emirates
Osprey (Pandion haliaetus) in flight catching a fish, Finland, July
Yellow-headed Picathartes / White necked rockfowl (Picathartes gymnocephalus) Kambui Hills, Sierra Leone, Vulnerable species. Photograph taken on location for BBC Africa series, September 2010
Wild female Amur leopard (Panthera pardus orientalis) on rocky hillside, Kedrovaya Pad reserve, Primorsky Krai, Far East Russia, January. Critically endangered species
Giant Panda cub (Ailuropoda melanoleuca) climbing. Yuan Meng, first Giant panda even born in FranceGiant Panda cub (Ailuropoda melanoleuca) climbing.Yuan Meng, first Giant panda even born in France, is now aged 8 months. and likes very much to stay in trees, Beauval Zoo, France