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RF - Bluestreak cleaner wrasse (Labroides dimidiatus) tending to Ribboned sweetlips (Plectorhinchus polytaenia). Yillet Kecil, Yillet Islands, Misool, Raja Ampat, West Papua, Indonesia
Polar bear (Ursus maritimus) mother and large second year cub swim across calm water, Scott Inlet, Baffin Island, Canadian Arctic, August. Vulnerable species
Hoffmanns Two-toed sloths (Choloepus hoffmanni) two orphan babies in Aviarios Sloth Sanctuary, Costa Rica
Two Konik Wild Horse (Equus ferus caballus) interacting. The Netherlands, November
RF- European edible frog (Rana esculenta) amongst duckweed, the Netherlands
Pair of Common toads (Bufo bufo) in amplexus underwater, Belgium, March
Head portrait of Satiny Parrot snake (Leptophis depressirostris) swallowing frog, Hungary, Europe
African Buffalo (Syncerus caffer) males fighting, Nakuru National park, Kenya
Blue Whales (Balaenoptera musculus) throat inflated, filter feeding at sea surface. California, USA. Sequence 1 of 2
Humpback Whale (Megaptera novaeangliae) mother and calf. Tonga, South Pacific, September
Ruff (Philomachus pugnax) two males displaying in territorial dispute, Liminka Finland June
White rhinoceros (Ceratotherium simum) mother and calf browsing. Entabeni Safari Conservancy, Limpopo region, Waterberg, South Africa, October
Two Red Foxes (Vulpes vulpes) fighting. The Netherlands. August
Polar bear (Ursus maritimus) sow and cub sliding on their backs over the pack ice, much like ice skating, 1002 coastal plain of the Arctic National Wildlife Refuge, Alaska, USA
Salmon landing on head of Grizzly bear (Ursus arctos horribilis) as it is leaping up rapids, Katmai National Park, Alaska, USA
Killer WHale / Orca (Orcinus orca) bull and cow in Resurrection Bay, Kenai Fjords National Park, outside Seward, Alaska, May
Great Egrets (Ardea alba) territorial fight above nest colony, Everglades National Park, Florida, USA, February
Common peafowl (Pavo cristatus) perched on trees at dawn, Western Ghats, Southern India
Goldfinches (Carduelis carduelis) squabbling near seed feeder, Berwickshire, Scotland, April
Two large white piglets (Sus scrofa domestica) at 3 weeks in sty, UK
Two Least Chipmunks (Tamias minimus) sitting on tree stump feeding, Yellowstone National Park, Wyoming, USA, October, crop of 1373882
Hanuman Langur / Northern Plains Grey Langur (Semnopithecus entellus) mother with baby in tree. Kanha National Park, Madhya Pradesh, India
Sperm Whale (Physeter macrocephalus) calf swimming uder its mother. Sao Miguel Island, Azores, July
Manta Ray (Manta Birostris) and Devil Ray (Mobula tarapacana) in background, accompanied by Pilot Fish (Naucrates ductor) and Remora. Santa Maria Island, Azores, September
Two Scarlet macaws (Ara macao) flying, captive
Ginger Kittens (Felis catus) in garden. France, September
Portrait of a Lynx (Lynx lynx) mother and kitten. Hanau, Germany, July. Captive
Snow goose with gosling (Chen caerulescens) at sunset. Taken on location for BBC Frozen Planet series, 2008
Indian / Asian one-horned rhinoceros (Rhinoceros unicornis) approaching, Kaziranga National Park, Assam, India
Yorkshire Terrier, with Netherland-cross rabbit
Young windmill-eared rabbit and matching guinea-pig
Two ginger kittens rolling playfully on their sides
Two ginger kittens lying on their sides
Black-and-white Border Collie bitch lying chin on floor with black-and-white kitten
Two male Mute swans (Cygnus olor) displaying on water in territorial dispute, UK
Two Denise's Pygmy Seahorses (Hippocampus denise) on a seafan (Muricella sp. ). Digital compositeTwo Denise's Pygmy Seahorses (Hippocampus denise) on a seafan (Muricella sp.). Digital composite. Misool, Raja Ampat, West Papua, Indonesia, November
Playful Border Collie puppies, 6 weeks
Border Collie with one of her puppies, 5 weeks, yawning
Narwhal (Monodon monoceros) crossing tusks above water surface. Baffin Island, Nunavut, Canada, April
Two orphan baby Elephants (Loxodonta africana) being affectionate. David Sheldrick Wildlife Trust Nairobi Elephant Nursery, Kenya, July 2010
Wild stallion (Equus caballus) lying down in the sagebrush and yawning. The foothills south of Reno, Nevada, USA, May
Female Rocky Mountain bighorn sheep ewe (Ovis canadensis canadensis) warning her newborn lamb that a rocky ledge is too close and dangerous
Two Brown bears (Ursus arctos) fighting in woodland wetlands, Kuhmo, Finland, July
Southern plains grey / Hanuman langur {Semnopithecus dussumieri} two adolescents sitting, Thar desert, Rajasthan, India
A young California sealion (Zalophus californianus) playing with a starfish, inside an underwater cave. The pups will pick up a variety of objects (such as starfish, lumps of coral)
Feral domestic horses (Equus caballus) wild herd stallion (grey) fighting a bachelor chestnut stallion, Blue Swallow Reserve, near Kaapsehoop, Mpumalanga province / Eastern Transvaal, South Africa
Two African lion cubs (Panthera leo) playing on a road in the Kalahari Desert, Botswana
European hare (Lepus europaeus) mating pair boxing in field, Slovakia