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Rock beauty angelfish (Holacanthus tricolor) pair at dusk performing their courtship dance with the larger male circling the female, Georgetown, Grand Cayman, Cayman Islands, British West Indies
Western grebe (Aechmophorus occidentalis) pair swimming, with reflections, Bolsa Chica Ecological Reserve, California, USA, February
Two Black-capped chickadees (Poecile atricapillus) in mid-air fight by a sunflower, New York, USA, February
Chilean huemul or South Andean deer (Hippocamelus bisulcus), Torres del Paine National Park, Patagonia, Chile. Endangered species
Adult male Hippopotamuses (Hippopotamus amphibius) posturing in agressive yawn behaviour, Luangwa River, South Luangwa National Park, Zambia, November. Digital composite (two frames stitched together)
African lions (Panthera leo) one lioness licking another, Okavango Delta, Botswana
Chinese Crested Dog, hairless bitch with puppy, male, 4 months
Male African Lion (Panthera leo) playing with 4 month cub. Big Marsh, near Ndutu, Nogorongoro Conservation Area, Serengeti National Park, Tanzania
Male Greenfinches (Carduelis chloris) squabbling in flight. Dorset, UK, March
Leaf cutter ant (Atta sp) female worker carrying leaf to nest, with a smaller minor ant riding. The minors job is to protect her sister from parasitic flies. Costa Rica, December
Green crowned brilliant hummingbird (Heliodoxa jacula) male taking off to feed at feeder in garden, another waiting perched on lichen covered branch, Costa Rica, January
Puffins (Fratercula arctica) pair take off together to go fishing. Farne Islands, Northumberland, UK, May. Non-ex
Pipevine Swallowtail Butterfly (Battus philenor) pair mating. Sinton, Corpus Christi, Coastal Bend, Texas, USA, June
A wild newborn colt (Equus caballus) standing up being nuzzled by its mother. Foothills of Reno, Nevada, USA, May
Domestic pig, free range sow and piglet sniffing noses, UK, August 2010
Dalmation dogs playing on beach, Norfolk, UK, August
Asiatic wild ass (Equus hemionus) with foal, captive, UK, endangered species
A pair of Toco Toucans (Ramphastos toco) flying across the Piquiri River at dawn, northern Pantanal, Mato Grosso, Brazil, September. (Digitally Modified)
A pair of Australian Sea Lions (Neophoca cinerea) socialising in the shallows. Kangaroo Island, South Australia, Australia. Southern Ocean. November
Two Cheetahs (Acinonyx jubatus) touching noses in greeting display, Serengeti NP, Tanzania, East Africa, January
Bobtail / Old English Sheepdog, sitting alongside a longhaired Chihuahua, tan and white coated
Great Dane, black coated bitch, and Magyar Vizsla / Hungarian Pointer, tan coated maleGreat Dane, black coated bitch, and Magyar Vizsla /Hungarian Pointer, tan coated male, double portrait in profile together
African Elephant (Loxodonta africana) bulls in dominance fight, Mono Pools national Park, Zimbabwe, Southern Africa
Roan antelope (Hippotragus equinus) with young offspring, Mlilwane nature reserve breeding programme, Swaziland, Southern Africa
Two Cape Ground squirrels (Xerus inauris) greeting, Kgalagadi Transfrontier Park, South Africa Non-ex
Polar bear (Ursus maritimus) sow with her cub outside their den in late winter, Arctic coast of Alaska
Wild horse / mustang (Equus caballus) two young mustangs play-fighting in the Pryor Mountains, Wyoming, USA
Trumpeter Swans (Cygnus buccinator) in winter morning mist, showing aggression during courtship behaviour, Mississippi River, Minnesota, USA, February
Snares-crested penguin (Eudyptes robustus) walking along rocks, Snares Island, New Zealand Sub-Antarctic Islands. Endemic and vulnerable species
RF- Gannets (Morus) displaying, Great Saltee, County Wexford, Ireland. May
RF - Head portrait of black Andalusian mare and stallion meeting for the first time in Spain, Europe
Koala (Phascolarctos cinereus) mother with joey aged eight months, Queensland, Australia, captive
Gannet (Morus bassanus) breeding pair during part of their elaborate courtship ritual. Saltee Islands, Republic of Ireland, May
African Lion (Panthera leo) cub reaches for a moment of intimacy with its father, Northern Tuli Game Reserve, Botswana
Giraffe (Giraffa camelopardalis) female bending down to calf, Okavango Delta, Botswana
Japanese macaque (Macaca fuscata fuscata) mother with rare white furred baby, Jigokudani Valley, Nagano Prefecture, Japan. June
Two Limon glass frogs (Sachatamia ilex) sitting on plant, one above the other, Canande, Esmeraldas
Bengal Tigers (Panthera tigris tigris) adult female drinking water, her juvenile cub beside her. Bandhavgarh National Park, India, April
RF - Percheron horse, two walking through snow, one black, one dappled grey. Alberta, Canada. February. (This image may be licensed either as rights managed or royalty free.)
Two wild Carneddau Ponies mutually grooming on the snow-covered slopes of Pen yr Ole Wen, overlooking Tryfan and Llyn Ogwen. Snowdonia National Park, Bethesda, Wales, UK. December, 2022
Two European goldfinches (Carduelis carduelis) fighting over food in snow, Wrocław, Lower Silesia, Poland. February
Whooper swans (Cygnus cygnus) pair engaged in courtship as snow falls, Myvatn lake, Iceland, March
Polar bear (Ursus maritimus) female with cub walking over snow with mountains in background, Svalbard, Norway. April
Two Shetland sheepdogs sitting and lying down in snow, portrait, Waterford, Connecticut, USA. December
Alpine ibex (Capra ibex) two adult males in mountain landscape. Alps, Aosta Valley, Gran Paradiso National Park, Italy. September
Great crested grebe (Podiceps cristatus) pair, courtship dance at dawn, Sierra de Grazalema Natural Park, Andalusia, Spain. April
Two Great egrets (Ardea alba) face to face against white sky, Lake Csaj, Pusztaszer, Hungary. Winner of the Portfolio category of the Terre Sauvage Nature Images Awards competition 2015
Two Grey wolves (Canis lupus) play fighting in snow, Finland. November
