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Two European grey wolves (Canis lupus) in woodland, captive, Norway, February
Dalmatian Pelican (Pelecanus crispus) portrait, Lake Kerkini, Greece. February. Highly honoured in the Birds Category of Natures Best Photography Windland Smith Rice International Awards Competition
White naped cranes (Grus vipio) in flight, Kyushu, Japan
Golden-crowned Sifaka or Tattersalls Sifaka (Propithecus tattersalli) climbing down tree in forests near Andranotsimaty, Daraina, north east Madagascar. Critically Endangered
Rufous elephant shrews (Elephantulus rufescens) Botswana
Two male Black grouse (Lyrurus tetrix) displaying at lek, Utajarvi, Finland, April
Male Panther Chameleons (Furcifer pardalis) fighting. Nosy Be, Madagascar. Photograph taken on location for BBC Wild Madagascar Series, January 2010
Impala (Aepyceros melampus) females, Masai-Mara Game Reserve, Kenya
Young Olive baboons (Papio hamadryas anubis) playing with one hanging from the others tail, Nakuru National Park, Kenya
Mature Bighorn Rams (Ovis canadensis) on high mountain pass. Jasper National Park, Alberta, Canada, October
Red-bellied Lemur (Eulemur rubriventer) pair, Ialasatra, Madagascar
Chimpanzee (Pan troglodytes) mother and 4 month infant resting in tropical forest, Western Uganda
Chimpanzee (Pan troglodytes) mother and baby age two and a half, tropical forest, Western Uganda
Red fox (Vulpes vulpes) cubs, Hertfordshire, England, UK, May. Did you know? There are 45 different subspecies of Red fox, and as well as many colour variations
Blue tits (Paris caereleus) two feeding on red berries, Uto Finland October
Blue Sheep (Pseudois nayaur) on hillside, Kekexili, Qinghai, China, January
Bengal Tigers (Panthera tigris) sub-adults, approximately 17-19 months old, playfighting. Endangered. Bandhavgarh National Park, India. Non-ex. Sequence 1 of 2
Bengal Tiger (Panthera tigris tigris) mother and cub, Pench National Park, Madhya Pradesh, India, taken on location for Tiger - Spy in the Jungle
Bengal Tigers (Panthera tigris tigris) resting in the shade national park, India, taken on location for Tiger - Spy in the Jungle March 2007
Starlings (Sturnus vulgaris), adults perched on branch in winter feeding on apple. Lorraine France. 3rd Prize in the Melvita Nature Images Awards competition 2013
Common darter dragonflies (Sympetrum striolatum) two females resting on reed, Dunsdon Nature Reserve, Devon, UK. August
RF- Two Grey wolves (Canis lupus) head portraits with damp coats from rain shower, captive
Razorbill (Alca torda) pair calling, between soft focus vegetation. Great Saltee, Saltee Islands, Co. Wexford, Ireland, June
Red foxes (Vulpes vulpes) fighting in snowy field with woodland in the background, Vosges, France, February
White stork (Ciconia ciconia) adult portrait, captive, Vogelpark Marlow, Germany, May
Japanese Macaques (Macaca fuscatata) grooming in hot springs, Jigokudani, Nagano Prefecture, Honshu, Japan
Two Guillemots (Uria aalge), Vardo, Norway, March
Newly hatched ducklings with eggs
Great crested grebe (Podiceps cristatus) pair performing courtship display at dawn, backlit with surrounding by mist, Cheshire, UK, March
RF- Gannet (Morus) in flight, Great Saltee, County Wexford, Ireland. April
Australian sealion with young at sea surface {Neophoca cinerea} Australia. Endangered species
Grizzly bears (Ursus arctos horribilis) looking for salmon, Brooks river, Katmai national park, Alaska, North America
Lion (Panthera leo) older cub playing with a younger one, Masai-Mara Game Reserve, Kenya. Vulnerable species
Two Galapagos brown pelicans (Pelecanus occidentalis) on beach, Floreana Island, Galapagos, October 2016
Two Common squirrel monkeys (Saimiri sciureus) amongst vegetation, Yasuni National Park, Orellana, Ecuador
Two Galapagos marine iguanas (Amblyrhynchus cristatus) on rock, Espanola Island, Galapagos, Vulnerable species
RF- Puffins (Fratercula) displaying. Skomer Island, Wales, UK. April
Greenfinch (Carduelis chloris) pair, one perched on branch and one hovering in snowfall, Derbyshire, February
Gannet (Morus bassanus) adult stretching its wings. Shetland Islands, Scotland, UK, September
Kingfisher (Alcedo atthis) flying up from water with caught fish, Worcestershire, UK, March
Pair of Tiger shrimps (Phyllognathia ceratophthalmus) on top of piece of coral. Bitung, North Sulawesi, Indonesia. Lembeh Strait, Molucca Sea
RF - Bluestreak cleaner wrasse (Labroides dimidiatus) tending to Ribboned sweetlips (Plectorhinchus polytaenia). Yillet Kecil, Yillet Islands, Misool, Raja Ampat, West Papua, Indonesia
Polar bear (Ursus maritimus) mother and large second year cub swim across calm water, Scott Inlet, Baffin Island, Canadian Arctic, August. Vulnerable species
Hoffmanns Two-toed sloths (Choloepus hoffmanni) two orphan babies in Aviarios Sloth Sanctuary, Costa Rica
Two Konik Wild Horse (Equus ferus caballus) interacting. The Netherlands, November
RF- European edible frog (Rana esculenta) amongst duckweed, the Netherlands
Pair of Common toads (Bufo bufo) in amplexus underwater, Belgium, March
Head portrait of Satiny Parrot snake (Leptophis depressirostris) swallowing frog, Hungary, Europe