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American pit bull, male, standing on grass and autumn leaves, portrait, Haddam, Connecticut, USA. October
Jack Russell terrier puppy standing on grass in autumn, portrait, Haddam, Connecticut, USA. October
Golden retriever standing at water's edge during spring thaw, Portage Creek, Portage, Alaska, USA. April
Yorkshire terrier standing among spring flowers, portrait, Haddam, Connecticut, USA. April
German shepherd dog, female, standing on rocks, portrait, Jacksonville, Oregon, USA. July
Whippet standing next to wooden fence and pink Roses, portrait, Haddam, Connecticut, USA. June
Grizzly bears (Ursus arctos) on a waterfall, waiting for leaping Salmon, Brooks Falls, Katmai National Park, Alaska, USA. July
Bengal kitten, aged 15 weeks, standing on hind legs reaching out, portrait
Common ostriches (Struthio camelus), male and female, standing at waterhole in front of Grant's zebras (Equus quagga boehmi). Tsavo East National Park, Kenya
Polar bear (Ursus maritimus) very young cub, probably only a few months old, standing up to take a look around, northern Svalbard, Norway, June
Dartmoor pony (Equus ferus caballus), endangered rare breed, mare and colt standing alert. Dartmoor National Park, Devon, England. June
Two Egyptian geese (Alopochen aegyptiaca) goslings standing side by side, London, UK. February
Anadolu stallion standing alert, portrait, Sirince mountains, Turkey
Two Komodo dragons (Varanus komodoensis) males, fighting on a remote beach, Komodo Island, Komodo National Park, Indonesia. Endangered
Two male Polar bears (Ursus maritimus) standing on hind legs, sparring, Churchill, Canada. November
Two Wolves (Canis lupus) standing in deep winter snow, Yellowstone National Park, USA. January
Putorana snow sheep (Ovis nivicola borealis) lamb, standing precariously on sheer cliff face, Putoransky State Nature Reserve, Putorana Plateau, Siberia, Russia
American Purple gallinule (Porphyrio martinica) balancing on a lily pad, Everglades National Park, Florida, USA
Dungeness crab (Metacarcinus magister) standing on seabed, Vancouver, British Columbia, Canada, Pacific Ocean
Black stork (Ciconia nigra), Great white heron (Ardea alba), Grey heron (Ardea cinerea) and Eurasian spoonbill (Platalea leucorodia) standing in shallow water, Pusztaszer reserve, Hungary. May
Grey heron (Ardea cinerea) standing at waters edge, Pusztaszer reserve, Hungary. May
Female Red fox (Vulpes vulpes) observing the yard from the top of the old stone wall, trying to determine whether it is safe to enter, Hungary. January
Grey heron (Ardea cinerea) on lake at sunset, Lake Csaj, Kiskunsagi National Park, Hungary. January
Male Ring ouzel (Turdus torquatus) standing on a garden lawn, Norfolk, UK. April
Matriarch Timber wolf (Canis lupus lycaon) sitting next to alpha male standing in snow. Hudson Bay, Manitoba, Canada
Emperor penguin (Aptenodytes forsteri) group on ice floe, Ross Sea, Antarctica. January
RF - Cheetah (Acinonyx jubatus) female patrolling territory. Long Gully, Ngorongoro Conservation Area, Tanzania. April
Mareeba rock wallaby (Petrogale mareeba) standing on granite boulders, Atherton Tablelands, Queensland, Australia
Male Numbat (Myrmecobius fasciatus) in dry woodland, Dryandra woodland, Western Australia
Dippers (Cinclus cinclus) standing on ice, ready to fight over feeding territory, Kuusamo, Finland, February
Caribbean flamingo (Phoenicopterus ruber) flock, Ria Lagartos Biosphere Reserve, Yucatan Peninsula, Mexico, June. Bookplate
Red fox (Vulpes vulpes) on snow covered frozen pond, Acadia National Park, Maine, USA. February
Fallow deer (Dama Dama) stags in velvet standing in a shady woodland spot, Knepp Estate, Sussex, UK, June
Adult White stork (Ciconia ciconia) standing beside three large chicks on their nest, Knepp Estate, Sussex, UK, June
Cheetahs (Acinonyx jubatus) on alert, Private reserve, Namibia, Africa. Captive
Wild boar (Sus scrofa) piglet. Forest of Dean, Gloucestershire, England, UK. March
Andean flamingo (Phoenicoparrus andinus) flock standing in water, Laguna Colorado, Eduardo Avaroa Andean Fauna National Reserve, altiplano of Bolivia. March
Arctic fox (Alopex lagopus) in snow. Svalbard, Norway. April
Polar bear (Ursus maritimus) male female pair looking upwards, snow covered hills in background. Svalbard, Norway, April 2018
Coyote (Canis latrans) standing near waters edge, in snow. Hayden Valley, Yellowstone National Park, USA. January 2019
King penguin (Aptenodytes patagonicus), two sleeping on snow. Grytviken, South Georgia. March
Snow leopard (Unica unica) standing beside boulder in rocky landscape of Himalayas, prior to scent marking. Ule, Ladakh, India. March 2018. Sequence 1/2
Pere Davids deer (Elaphurus davidianus) two stags boxing amongst herd of hinds. Captive bred in reserve after reintroduction from Woburn Abbey, England, UK
Amur / Siberian tiger (Panthera tigris altaica) pair in courtship, in snow. Captive in tiger park, Heilongjiang Province, China. January
RF- Reindeer (Rangifer tarandus) feeding. Denali National Park, Alaska, USA
Reticulated Giraffe (Giraffa camelopardalis reticulata) standing in grassland. Laikipia, Kenya. October
