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Redwing (Turdus iliacus) in winter on windfall apples. Dorset, UK, December 2010
Tibetan antelope (Pantholops hodgsonii) male, Kekexili, Qinghai, Tibetan Plateau, China, November
Two Irish Gypsy cob mares (Equus caballus), one dapple grey - grazing and one piebald - walking on rough pastureland on Hackpen Hill with a Beech clump (Fagus sylvaticus) in the background
Male Ostrich (Struthio camelus) protecting chicks from the sun with its wings, Samburu National Reserve, Kenya, Africa
Female Grizzly bear (Ursus arctos horribilis) standing up, with two cubs nearby, Khutzeymateen Grizzly Bear Sanctuary, British Columbia, Canada, June
Lesser flamingo (Phoeniconaias minor) portrait, Lake Bogoria, Rift valley, Kenya, Africa
Topi (Damaliscus korrigum) at sunset, Masai-Mara Game Reserve, Kenya
Red deer (Cervus elaphus) stag in autumn woodlands, Bradgate Park, Leicestershire, UK, October
RF - European Badger (Meles meles) foraging at tree trunk in deciduous woodland. Mid Devon, UK. June
Red deer (Cervus elaphus) stag in the snow, Scotland, March
Puffin (Fratercula arctica) standing on Sea thrift (Armeria maritima) with wings out stretched with a beak full of sand eels. Close-up portrait. Fair Island, Shetland Islands, Scotland, July
RF- Giraffe silhouette at sunset. Namibia, Etosha National Park
Leopard (Panthera pardus) hunting Springbok (Antidorcas marsupialis) Etosha, Namibia, Finalist in the Mammals Category of the Wildlife Photographer of the Year 2015
Forest reindeer, (Rangifer tarandus fennicus) in velvet, Viiksimo, Kuhmo region. Finland, July. Rare subspecies which were nearly extinct in the 19th Century
RF- Alpine marmot (Marmota marmota) reaching upwards, Hohe Tauern National Park, Austria. July
Gemsbok (Oryx gazella) female with two calves, Namibrand Reserve, Namib Desert, Namibia, Nambia
Great white egret (Ardea alba) splashing, Hungary, January
Young Meerkat (Suricata suricatta) standing on its hind legs near the edge of Makgadikgadi Pans National Park, Botswana, April
Male Caucasian turs (Capra caucasica) rutting with Dykh-tau Mountain in the background, Kabardino-Balkarsky Zapovednik / Reserve, Russia, July
Lowchen / Little Lion dog, black and white, partially shaved, portrait in garden, France
Chinese Crested Dog, hairless
Five newly hatched chicks in a row
Four Wisent / Bison (Bison bonasus) standing in a row. Bialowieza Forest, Bialowieza National Park, Poland, November
Siamese kitten, 10 weeks, reaching up
Tabby kitten, 5 months, standing
Portrait of a silver tabby-and-white cat walking
Moose (Alces alces) head portrait of bull standing in forest clearing, Cap Breton Highlands National Park, Nova Scotia, Canada, September
Yellow labrador, five puppies in a row
Miniature Schnauzer, black-silver coated, standing in autumn foliage
Pygmy x Golden Guernsey female goat kid, standing on hind legs
King Penguin (Aptenodytes patagonicus) adult surrounded by huddled chicks, riding out a snowstorm at Gold Harbor, South Georgia
Polar bear (Ursus maritimus) female and her two cubs on large iceberg off Baffin Island. Nunavut, Canada, August 2010
Meerkat / Suricate family group (Suricatta suricata) standing alert together, Kalahari Meerkat Project, Van Zylsrus, Northern Cape, South Africa
Wild Horse / Mustang pinto stallion rearing, McCullough Peaks Herd Area, Cody, Wyoming, USA
RF- Masai giraffe (Giraffa camelopardalis tippelskirchi) two young calves together, Masai Mara Game Reserve Kenya
Polar bear (Ursus maritimus) curious cub sits up on its hind legs, paw raised, and tries to balance itself, along Bernard Spit in autumn, 1002 area of the Arctic National Wildlife Refuge, Alaska
Baby African elephants (Loxodonta africana) of various ages playing together, Addo national park, Eastern Cape, South Africa, January
Female Cheetah (Acinonyx jubatus) standing in long grass on the savannah, Okavango Delta, Botswana, Africa
Black Grouse (Tetrao tetrix) males at lek, Utajarvi, Finland, April
RF- Whooper Swan (Cygnus cygnus) standing on frozen loch at dawn in winter. Loch Insh, Cairngorms National Park, Scotland, UK
American pine marten (Martes americana) standing on hind legs in deep snow, Yellowstone National Park, USA. January
Iberian ibex (Capra pyrenaica), adult male, in snow covered landscape after snowfall. Sierra Nevada National Park, Andalusia, Spain. February
Collie standing in snowy, frozen marsh, Eagle River, Alaska, USA. April
Alpine ibex (Capra ibex) two adult males in mountain landscape. Alps, Aosta Valley, Gran Paradiso National Park, Italy. September
Weasel (Mustela erminea) in white winter coat standing upright in snow, Upper Bavaria, Germany, Europe. February
Startled Eurasian lynx (Lynx lynx) kitten, aged eight weeks, breeding and reintroduction program, Germany. Captive. August
Red fox in a snow storm (Vulpes vulpes) Winter winds on Kamchatka can reach 40 meters a second. Kronotskty Zapovednik, Kamchatka, Russia
Mandarin duck (Aix galericulata) drake standing on frozen pond during snowstorm. Richmond Park, London, UK. January