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Bharal / Blue sheep (Pseudois nayaur) herd standing on a rocky slope, higher Himalaya mountains, Kibber Wildlife Sanctuary, India. March
Guanaco (Lama guanicoe) in grassland with Torres del Paine rock towers in background. Torres del Paine National Park, Patagonia, Chile. January 2020
Cuban hutia (Capromys pilorides) stands up on its back legs on a tropical beach. Jardines de la Reina, Gardens of the Queen National Park, Cuba. Endemic
Mustang herd standing in grassland. Red Desert Complex, Wyoming, USA. September
Grey squirrel (Sciurus carolinensis) stood upright on short grass. London, England, UK. March
European Brown Bear (Ursus arctos) Cubs standing amongst cotton grass, Finland, June
Roe deer (Capreolus capreolus) buck in woodland. Peerdsbos, Brasschaat, Belgium. August
Carters semaphore gecko (Pristurus carteri) standing on rocky ground. Oman, June
RF - American bison (Bison bison) male standing near Punch Bowl thermal pool, green algae in foreground. Yellowstone National Park, USA
African penguin (Spheniscus demersus) standing on rock at dusk, with clustered penguins in background. South Africa. November
Arctic fox (Alopex lagopus) standing in tundra. Svalbard, Norway, July
White-tailed deer (Odocoileus virginianus) doe and fawn standing amongst Bulrushes on snow covered pond. Acadia National Park, Maine, USA. January
RF - Guanaco (Lama guanicoe) standing in front of mountain towers of Paine. Torres del Paine National Park, Patagonia, Chile. December 2018
Puma (Puma concolor puma) young male standing with mouth open. Estancia Amarga, near Torres del Paine National Park, Patagonia, Chile. December
American bison (Bison bison), two females covered in hoar frost near hot spring, portrait. Midway Geyser Basin, Yellowstone National Park, USA. February
Guanaco (Lama guanicoe) standing, towers of Torres del Paine National Park in background. Patagonia, Chile. December 2018
American quarter horse, three standing in snow. Alberta, Canada. February
Sorraia horses, mare and foal standing amongst wildflowers in meadow. Middle Coa, Coa Valley, Western Iberia, Portugal. April
Wild cat (Felis sylvestris) in snow, Vosges, France, February
Northern white cheeked gibbon (Nomascus leucogenys) female standing in tree with baby aged two weeks. Parents captive bred and released
Little penguin (Eudyptula minor) standing on rocks of St Kilda breakwater, returning to nest at dusk. Melbourne city in background. Victoria, Australia. December 2016
RF- Fallow deer (Dama dama) doe at sunrise during rut, Deer Park, Holkham, Norfolk, UK, October
Blue-footed booby (Sula nebouxii) colony at sunet. Cape Douglas, Fernandina Island, Galapagos
People watching the Fissure eruption of Chico volcano into 9km diameter caldera. Sierra Negra volcano, Isabela Island, Galapagos, Ecuador. October 2005
Eurasian otter (Lutra lutra) feeding whilst on log in woodland burn, Glenfeshie, Cairngorms National Park, Scotland, UK, September
Red squirrel (Sciurus vulgaris) standing on log surrounded by heather, Cairngorms National Park, Scotland, UK, August
Dipper (Cinclus cinclus) standing on ice at waters edge, at sunset. Kuusamo, Northern Ostrobothnia, Finland. December
Blue-point kitten playing, standing on hind legs
Tabby kitten, age 6 weeks, standing with raised paws
Red fox (Vulpes vulpes) cub looking over shoulder at camera. Sheffield, England, UK. May
Red fox (Vulpes vulpes) cub looking at camera, in morning. Sheffield, England, UK. June
Turtle dove (Streptopelia turtur) standing at a drinking pool. Hungary. May
Muskox (Ovibos moschatus) in tundra with mountains in background, Dovrefjell National Park, Norway. September 2018
Thomsons gazelle (Eudorcas thomsonii) herd. Serengeti National Park, Serengeti, Tanzania
Thomsons gazelle (Eudorcas thomsonii) male and two females standing in rainstorm, Masai Mara Game Reserve, Kenya
Great white egrets (Casmerodius albus) reflected in Pulicat Lake, Tamil Nadu, India, January 2013
Newfoundland dog, male standing in snow. Killingworth, Connecticut, USA
Hoatzin (Opisthocomus hoazin), pair on floating log in Cocha Salvador ox-bow lake. Manu Biosphere Reserve, Peru
A Kamchatka Brown Bear (Ursus arctos beringianus) standing in breaking waves. Pacific coast of Kronotsky Zapovednik Nature Reserve, Kamchatka Peninsula, Russian Far East, July
African penguin (Spheniscus demersus) returning to colony. Near Simons Town in False Bay, between Fish Hoek and Cape Point, Western Cape, South Africa. Endangered species
Ginger and white kitten looking at camera
Wild rare Losino stallion standing in snow, Losa Valley, Burgos, Spain. January 2014
Bay Icelandic horse feeding in the snow, Snaefellsnes Peninsula, Iceland, March
Wild Konik horses with Starlings (Sturnus vulgaris) Odry delta reserve, Stepnica, Poland, July
Meerkat (Suricata suricatta) standing and keeping watch while another excavates a burrow in soft sand before disappearing underground for the night. Makgadikgadi Pans, Botswana, April
Cheetah (Acinonyx jubatus) female standing on mound to look for a prey, Masai Mara Game Reserve, Kenya, September
Western sandpipers (Calidris mauri) roosting on barrier island tidal flats. Terrebonne Parish, Louisiana, USA. October
Eurasian brown bear (Ursus arctos) rubbing back against tree, Suomussalmi, Finland, July 2008