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Arctic fox (Vulpes lagopus) resting in grass, Isfjorden, Svalbard, Norway, July
Leatherback turtle (Dermochelys coriacea) female, returning to sea after laying eggs in nest on beach, Grande Riviere, Trinidad Island, Trinidad & Tobago, Caribbean
Grey reef shark (Carcharhinus amblyrhynchos) swimming, Yap, Micronesia, Pacific Ocean
Nesting female Olive ridley turtle (Lepidochelys olivacea) passing new hatchling. Playa Escobilla Sanctuary, Oaxaca, Mexico
Asiatic lion (Panthera Leo persica) female, portrait, Gir National Park, Gujarat, India
Leopard (Panthera pardus) standing in grassland, India
Adult Common tern (Sterna hirundo) brooding two chicks. Long Island, New York, USA. June
Male Downy woodpecker (Picoides pubescens) taking off to avoid Male Hairy woodpecker (Picoides villosus) that landed on its perch in winter, with orange foliage in background
Small pearl-bordered fritillary (Boloria selene) roosting on ribwort plantain (Plantago lanceolata), Marsland mouth, Devonshire, UK. May
Small white butterfly (Pieris rapae) resting on bluebell (Hyacinthoides non-scripta), Boscastle, Cornwall, UK. April
Silver studded blue butterfly (Plebejus argus) resting on grass blade, Gwithian Towans, Cornwall, UK. June
Marbled white butterfly (Melanargia galathea) resting on common spotted orchid (Dactylorhiza fuchsii), Dunsdon, Devon, UK. July
Osprey (Pandion haliaetus) fledgling strengthening wing in nest. Rasa Island Special Biosphere Reserve, Sea of Cortez, Mexico. April
Springbok (Antidorcas marsupialis) herd in grassland after rain, Palmwag Conservancy, Damaraland, Namibia
Gemsbok (Oryx gazella) pair standing in grass after wet season in desert, Namib-Naukluft National Park, Namibia
Gemsbok (Oryx gazella) female and calf walking past sand dune, Sossusvlei UNESCO World Heritage Site, Namib desert, Namib-Naukluft National Park, Namibia
Humpback whales (Megaptera novaeangliae) pair engaging in courtship behavior, dancing for hours, off Tubuai, French Polynesia, Pacific Ocean
Sika deer (Cervus nippon) doe standing on rocky outcrop overlooking mountain forest, Land of the Leopard National Park, Russian Far East. Taken with remote camera. January
Siberian tiger (Panthera tigris altaica) sitting and scent marking tree in forest by rubbing body against it, Land of the Leopard National Park, Russian Far East. Endangered
Amur leopard (Panthera pardus orientalis) walking past snow-covered trees on mountain slope, Land of the Leopard National Park, Russian Far East. Critically endangered. Taken with remote camera
Australian shepherd standing on frozen pond, Eagle River, Alaska, USA. April
Siberian husky standing on rock in autumnal woodland, New Hampshire, USA. October
Golden retriever standing at water's edge during spring thaw, Portage Creek, Portage, Alaska, USA. April
German shepherd dog, female, standing on rocks, portrait, Jacksonville, Oregon, USA. July
Whippet standing next to wooden fence and pink Roses, portrait, Haddam, Connecticut, USA. June
Grey seal (Halichoerus grypus) pup and mother sniffing and nuzzling each other. Donna Nook, Lincolnshire, UK. November
Humpback whale (Megaptera novaeangliae) surfacing, Tubuai, French Polynesia, Pacific Ocean
Humpback whale (Megaptera novaeangliae) swimming just below the ocean surface, Tubuai, French Polynesia, Pacific Ocean
Humpback whale (Megaptera novaeangliae) creating bubble arc with fin after surfacing, Tubuai, French Polynesia, Pacific Ocean
Humpback whale (Megaptera novaeangliae) pair swim together, Tubuai, French Polynesia, Pacific Ocean
Pacific trumpetfish (Aulostomus chinensis) juvenile swimming near the water surface in the open ocean, Tubuai, French Polynesia, Pacific Ocean
Humpback whale (Megaptera novaeangliae) creating bubble arc with fin at waters surface, Tubuai, French Polynesia, Pacific Ocean
Humpback whale (Megaptera novaeangliae) swimming towards surface, Tubuai, French Polynesia, Pacific Ocean
Humpback whale (Megaptera novaeangliae) swimming below water surface with snorkeler in the background, Tubuai, French Polynesia, Pacific Ocean
Common brimstone butterfly (Gonepteryx rhamni) flying towards Purple betony (Betonica officinalis) flowers, Upper Bavaria, Germany. July
Lion (Panthera leo) male, juvenile, resting in tree during middle of the day to escape the heat, near Ndutu, Ngorongoro Conservation Area / Serengeti National Park border, Tanzania
Lion (Panthera leo) female, resting in tree during middle of day to escape the heat with juvenile male Lion approaching, near Ndutu, Ngorongoro Conservation Area / Serengeti National Park border
Lion (Panthera leo) male, scent marking its territory at dawn, Ngorongoro Conservation Area / Serengeti National Park boundary, northern Tanzania
Puma (Puma concolor) female, walking in deep snow, Torres del Paine National Park / Estancia Laguna Armarga, Patagonia, Chile
Guanaco (Lama guanicoe) herd with the Towers granite rock formation and low cloud in background, Torres del Paine National Park, Patagonia, Chile
Puma (Puma concolor) female, cleaning paw of compacted snow, Torres del Paine National Park / Estancia Laguna Armarga, Patagonia, Chile
Puma (Puma concolor) female, on snowy hillside, Torres del Paine National Park / Estancia Laguna Armarga, Patagonia, Chile
Three Guanacos (Lama guanicoe) running over hillside in front of the Towers granite rock formation and low cloud, Torres del Paine National Park, Patagonia, Chile
Puma (Puma concolor) female, running sequence in deep snow, Torres del Paine National Park / Estancia Laguna Armarga, Patagonia, Chile. Composite of five separate images
Puma (Puma concolor) female, with frozen whiskers, head portrait, Torres del Paine National Park / Estancia Laguna Armarga, Patagonia, Chile
Guanaco (Lama guanicoe) chewing grass, head portrait, Torres del Paine National Park, Patagonia, Chile
European common blue (Polyommatus icarus) male resting on a flower, Estonia, June
Grizzly bears (Ursus arctos) on a waterfall, waiting for leaping Salmon, Brooks Falls, Katmai National Park, Alaska, USA. July