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Pectoral sandpiper (Calidris melanotos) juvenile feeding around edge of shallow flood, Llanrhystud, Ceredigion, West Wales, UK, September
Water vole (Arvicola amphibius) Kent, UK September
Red Squirrel (Sciurus vulgaris), at woodland pool, Cairngorms National Park, Scotland, UK.November
White rhinoceros (Ceratotherium simum) male and female, face to face, Mosi Oa Tunya National Park, Zambia
White rhinoceros (Ceratotherium simum) with a long horn, Marakele National Park, South Africa
Lesser spotted catshark / Dogfish shark (Scyliorhinus canicula) amongst seaweed on a maerl bed, South Arran Marine Protected Area, Isle of Arran, Scotland, UK, August
Brown hare (Lepus europaeus) adult in arable field, Scotland, August
Red squirrel (Sciurus vulgaris) standing on log surrounded by heather, Cairngorms National Park, Scotland, UK, August
Meadow Brown Butterfly (Maniola jurtina) feeding from Scabious flower (Scabiosa) North Downs, Surrey, England, UK, August
Clouded yellow butterfly (Colias croceus) on Fleabane. Dorset, UK, August
Brimstone butterfly (Gonepteryx rhamni) on Devil s-bit Scabious (Succisa pratensis). Peak District National Park, Derbyshire, UK. August
California sealion (Zalophus californianus) swimming. Magdalena Bay, Baja California Sur, Pacific Ocean, Mexico
Striped marlin (Tetrapturus audax) feeding on Sardine (Sardinops sagax). Magdalena Bay, Baja California Sur, Pacific Ocean, Mexico
Dunlin (Calidris alpina) portrait. Vardo, Finnmark, Norway. May
Black woodpecker (Dryocopus martius) male perched on tree trunk, trees covered in hoar frost. Kuusamo, Northern Ostrobothnia, Finland. February
Dipper (Cinclus cinclus) standing on ice at waters edge, at sunset. Kuusamo, Northern Ostrobothnia, Finland. December
Hazel grouse (Tetrastes bonasia) standing in snow at forest edge. Helsinki, Finland. February
Colombian red howler monkey (Alouatta seniculus) mother and baby in tree. Northern Colombia
Pileated Woodpecker (Dryocopus pileatus) male, at feeding excavation in tree trunk in winter, Freeville, New York, USA, January
American oystercatchers (Haematopus palliatus) courting pair performing Piping Display, calling as they walk along together, Fort DeSoto Park, Florida, USA, March
Male Red winged blackbird (Agelaius phoeniceus) taking off, New York, USA, May
Galapagos green turtle (Chelonia mydas agassizi) swimming in coastal waters of San Cristobal Island, Galapagos
Red squirrel (Sciurus vulgaris) feeding in snow. Scotland, UK. March
Rivolis hummingbird (Eugenes fulgens) nectaring on Fuchsia (Fuchsia sp) flower. Costa Rica
Leaf-nosed bat (Phyllostomidae sp) flying towards Banana (Musa sp) flower to feed. Costa Rica
Green-crowned brilliant (Heliodoxa jacula), three feeding on Lobster-claw (Heliconia sp) flower in rain. Costa Rica
Red fox (Vulpes vulpes) cub looking over shoulder at camera. Sheffield, England, UK. May
Red fox (Vulpes vulpes) cub looking at camera, in morning. Sheffield, England, UK. June
Badger (Meles meles) eating apples in urban garden. Sheffield, England, UK. October
Turtle dove (Streptopelia turtur) standing at a drinking pool. Hungary. May
Hedgehog (Erinaceus europaeus) walking through puddle. Sado Estuary, Portugal. March
Mountain gorilla (Gorilla beringei beringei), dominant silverback in forest clearing. Volcanoes National Park, Rwanda
Muskox (Ovibos moschatus) in tundra with mountains in background, Dovrefjell National Park, Norway. September 2018
Flightless cormorant (Phalacrocorax harrisi), adult and two chicks. Beagle Crater, Isabela Island, Galapagos
Southern lechwe (Kobus leche), two males about to fight, facing each other. Okavango Delta, Botswana
Bamboo lemur (Hapalmur griseus) sitting on tree stump. Andasibe-Mantadia National Park, Madagascar
Thomsons gazelle (Eudorcas thomsonii) herd. Serengeti National Park, Serengeti, Tanzania
Thomsons gazelle (Eudorcas thomsonii) male and two females standing in rainstorm, Masai Mara Game Reserve, Kenya
Killer whale (Orcinus orca) hunting inside herring baitball (Clupea harengus), Andenes, Andoya island, North Atlantic Ocean, Norway, April
Adult Kea (Nestor notabilis) close-up portrait. Arthurs Pass National Park, New Zealand, July. Endangered Species
Madagascar / Malagasy kingfisher (Alcedo vintsioides) perching, Anjajavy Private Reserve, north west Madagascar
Kori bustard (Ardeotis kori) male courtship display, Kgalagadi Transfrontier Park, Northern Cape, South Africa
Seychelles Sunbird (Cinnyris dussumieri), adult male singing, Praslin Island, Republic of Seychelles
Komodo dragon (Varanus komodoensis) Rinca Island, Indonesia
Chamois (Rupicapra rupicapra) in the snow, Jura Switzerland, November
Thorny dragon (Moloch horridus) in desert habitat, Australia
Head portrait of black Andalusian stallion and mare meeting for the first time in southern Spain, Europe
Black korhaan (Eupodotis afra) male displaying and calling. Etosha National Park, Namibia, March
