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Pine marten (Martes martes) foraging in Pine (Pinus sp) woodland at night. Glenfeshie, Cairngorms National Park, Scotland, UK
Osprey (Pandion haliaetus) fishing. Rothiemurchus, Cairngorms National Park, Scotland, UK
Bigeye trevally (Caranx sexfasciatus), a large shoal. Cocos Island National Park, Costa Rica
Eurasian badger (Meles meles), two foraging in Pine (Pinus sp) woodland. Glenfeshie, Cairngorms National Park, Scotland, UK
Marine iguana (Amblyrhynchus cristatus) basking on volcanic rock at coast. Isabela Island, Galapagos
Silky shark (Carcharhinus falciformis). Cocos Island National Park, Costa Rica
Rufous motmot (Baryphthengus martii) perched on branch. Rancho Naturalista, Cartago, Costa Rica
Parrot snake (Leptophis ahaetulla) in aggressive pose with mouth open. Costa Rica
Keel-billed toucan (Ramphastos sulfuratus) perched on branch with beak open. Laguna del Lagarto, Alajuela, Costa Rica
Badger (Meles meles) emerging through gateway in wall at night. In urban area, Sheffield, England, UK. October 2018
Violet sabrewing hummingbird (Campylopterus hemileucurus) nectaring on Orchid. Costa Rica
Muskox (Ovibos moschatus), male portrait. Dovrefjell National Park, Norway
Red-footed booby (Sula sula), adult and chick at nest. Genovesa Island, Galapagos
Nazca booby (Sula granti) at nest with egg. Overlooking sea, Gardner Islet, Floreana Island, Galapagos. December 2014
Nazca booby (Sula granti), pair amongst Prickly pear (Opuntia sp) cacti at coast. Wolf Island, Galapagos. August 2016
Nazca booby (Sula granti), pair billing. Genovesa Island, Galapagos. May 2015
Lapwing (Vanellus vanellus) portrait in upland hay meadow, Upper Teesdale, CO Durham, England, UK, June
Australian sea lion (Neophoca cinerea), two females greeting each other on beach. Seal Bay Conservation Park, Kangaroo Island, South Australia
Great bowerbird (Chlamydera nuchalis), male showing off treasures whilst female inspects bower. Lake Argyle, Kununurra, Western Australia
Australian green tree frog (Litoria caerulea) camouflaged on Palm leaf. Lake Argyle, Kununurra, Western Australia
Melanistic leopard / Black panther (Panthera pardus) Nagarahole National Park, Nilgiri Biosphere Reserve, Karnataka, India
Green iguana (Iguana iguana) rear view of basking in sun, Banco Chinchorro Biosphere Reserve, Caribbean region, Mexico, May
Skeleton of a seahorse (Hippocampus sp. ) showing the remarkable structure of bonesSkeleton of a seahorse (Hippocampus sp.) showing the remarkable structure of bones that form a virtual exoskeleton, with few deep internal bones, in this unusual vertebrate group
Atlantic puffin (Fratercula arctica) portrait with beak open / calling. Shetland IslandsAtlantic puffin (Fratercula arctica) portrait with beak open/calling. Shetland Islands, Scotland, July
Black grouse (Lyrurus tetrix), male at lek, Finland, April
Kingfisher (Alcedo atthis) male, after diving, taking off from water with fish, a Common Roach (Rutilus rutilus) Lorraine, France, August
Zebra (Equus quagga) leaping during stampede, Serengeti, Tanzania. Vignette added and right edge expanded
Fin whale (Balaenoptera physalus) just below surface, south Barcelona coast, Spain, Mediterranean Sea
Fin whale (Balaenoptera physalus) with ctenophore in front of its mouth, south Barcelona coast, Spain, Mediterranean Sea
Proboscis monkey (Nasalis larvatus) mother suckling infant, Sabah, Malaysia
Harpy eagle (Harpia harpyja) close up head portrait, captive
Deccan ground gecko (Cyrtodactylus albofasciatus) on forest floor, backlit with water droplets. Amboli, Maharashtra, India. Endemic to Western Ghats
Strawberry poison frog (Oophaga pumilio) photographed on a white background in mobile field studio, Central Caribbean foothills, Costa Rica
Sperm whale (Physeter macrocephalus) resting just beneath surface, Faial Island, Azores, Portugal, Atlantic Ocean July
Short-finned pilot whales (Globicephala macrorhynchus) two just below surface, Los Gigantes, South Tenerife, Canary Islands, Atlantic Ocean, July
Killer whale (Orcinus orca) diving and hunting for herring fish (Clupea harengus), Andenes, Andoya island, North Atlantic Ocean, Norway, April
Great barracuda (Sphyraena barracuda) with three parasitic Copepods at end of upper jaw. Hawaii
Andean cock-of-the-rock (Rupicola peruvianus), two males perched in tree at lek in mid-altitude montane rainforest. Manu Biosphere Reserve, Amazonia, Peru
Chesapeake Bay retriever male standing in saltmarsh, Long Island Sound, Madison, Connecticut, USA
English springer spaniel in Rhododendron. Haddam, Middlesex, Connecticut, USA
Weimaraner in front of Rhododendron flowers, Haddam, Connecticut, USA. May
Parson's chameleon (Calumma parsonii), male hunting in forest canopy. Mitsinjo Forest, Andasibe-Mantadia National Park, eastern Madagascar
Bronzed frog / Common Wood Frog (Hylarana temporalis) Sri Lanka
Red fox (Vulpes vulpes) visiting woodland stream to drink at night. Camera trap image. Near Bath, England, UK. June
Collared peccary (Tayassu tajacu), adult drinking, South Texas, USA. May
Tortoiseshell-tabby kitten, age 10 weeks, walking across
Northern raven (Corvus corax) pair perching on branch. Danube Delta, Romania, May