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A female (in front) and male Sockeye salmon (Oncorhynchus nerka) over eggs in their spawning river. Adams River, British Columbia, Canada, October
Male Great white shark (Carcharodon carcharias) with sunrays, Guadalupe Island, Mexico, Pacific Ocean
Orange Tip Butterfly (Anthocharis cardamines) male resting on Bluebell (Endymion / Hyacinthoides nonscriptus). Peak DIstrict National Park, Derbyshire, UK. April
Toco Toucan (Ramphastos toco) head and beak profile portrait, Brazil
Marbled White butterfly (Melanargia galathea) resting on grass, Dunsdon Nature Reserve, Devon, UK. July
Leopard (Panthera pardus) hunting Springbok (Antidorcas marsupialis) Etosha, Namibia, Finalist in the Mammals Category of the Wildlife Photographer of the Year 2015
Marine iguana (Amblyrhynchus cristatus) underwater. Fernandina Island. Galapagos, Endemic Species
Blacktip reef shark (Carcharhinus melanopterus) juvenile in shallow water, Maldives, Indian Ocean
Scarlet ibis (Eudocimus ruber), perched, Coro, Venezuela
Great frigatebird (Fregata minor) male displaying inflated gular sac, Galapagos, Ecuador
Great white egret (Ardea alba) splashing, Hungary, January
Green / Double-crested basilisk (Basiliscus plumifrons) running across water surface, Santa Rita, Costa Rica
Svalbard reindeer (Rangifer tarandus platyrhynchus) walking in golden light. Svalbard, Norway. April
Great Blue shark (Prionace glauca) profile portrait viewed from slightly above, Pico Island, Azores, Portugal, Atlantic Ocean
Young Meerkat (Suricata suricatta) standing on its hind legs near the edge of Makgadikgadi Pans National Park, Botswana, April
Clouded yellow butterfly (Colias crocea) On Wild sweet pea flower, Captive, UK, July
Siberian forest cat kitten walking, 7 weeks, white coat
Wild Horse / Mustang shaking head and mane, Adobe Town Herd Area, southwestern Wyoming, USA, WINNER OF WILD HORSE SECTION, EQUINE IDEAL PHOTOGRAPHY CONTEST 2012
Dall sheep (Ovis dalli) yearling lamb running across alpine tundra, Denali National Park, Alaska, USA
Chinese Crested Dog, hairless
Wide-armed mantis (Cilnia humeralis) reaching out, photographed on a white background, originating from Africa. Captive
Tabby kitten, 5 months, standing
Mute Swan (Cygnus olor) with its beak in water. Fife, Scotland, November
A large female Tiger Shark (Galeocerdo cuvier) swimming in open water. Grand Bahama, Bahamas. Tropical West Atlantic Ocean. March
Free diver coming up from dive to explore coral reef, Sinai, Egypt, Red Sea Model released
European praying mantis (Mantis religiosa) camouflaged on leaf, Tuscany, Italy
Juvenile Green turtle (Chelonia mydas) swimming near the surface, split level view, Fakarava atoll lagoon, Tuamotu Archipelago, French Polynesia, Pacific Ocean
Red eyed tree frog (Agalychnis callidryas) La Selva Field Station, Costa Rica
Strawberry poison dart frog (Oophaga pumilio) La Selva Field Station, Costa Rica
Great egret (Ardea alba) flying across lake, Ranthambhore National Park, India
Ribbon sawtail fish (Idiacanthus fasciola) from Atlantic Ocean, at a depth of 800-1000m
Frilled shark (Chlamydoselachus anguineus) specimen with mouth open, from Atlantic Ocean, at a depth of 729m
Lanternfish (Lepidophanes guentheri) - deepsea species showing bioluminescence
Polar Bear (Ursus maritimus) sticking tongue out, Svalbard, Norway. July
Toco Toucan (Ramphastos toco) beak open with tongue visible while feeding on mango, Brazil
RF - African elephant (Loxodonta africana) calf walking with trunk raised, Amboseli National Park, Kenya, August
RF- Masai giraffe (Giraffa camelopardalis tippelskirchi) two young calves together, Masai Mara Game Reserve Kenya
Loggerhead sea turtle (Caretta caretta) emerging from shell, Dalyan delta, Turkey, July
Moose (Alces alces) bull portrait, Baxter State Park, Maine, USA
Arctic tern (Sterna paradisaea) in flight with fish in beak for feeding chick, Machias Seal Island, Bay of Fundy, New Brunswick, Canada, July
Crowned dwarf / Smooth-fronted caiman (Paleosuchus trigonatus) with mouth open, Cuyabeno, Sucumbios, Ecuador
Rio pescado harlequin toad (Atelopus balios) pair on branch, female on the left, Azuay, Ecuador, Critically endangered species, possibly extinct
Sword billed hummingbird (Ensifera ensifera) in flight, Guango, Napo, Ecuador
Juvenile Sumatran tiger (Panthera tigris sumatrae), aged four months, captive, occurs in Sumatra, Indonesia
RF- Young male Bengal tiger, head portrait in profile (Panthera tigris tigris). Bandhavgarh National Park, India
Spinner dolphin (Stenella longirostris) with possible calf, Kona coast, Hawaii, USA, August
Kori bustard (Ardeotis kori) male, performing mating display, silhouetted at dusk, El Karama Wilderness Conservancy, northern Kenya
Tufted duck (Aythya fuligula) on water on a misty morning, Valkenhorst Nature Reserve, Valkenswaard, The Netherlands. June