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Feral domestic cattle, Puerto Fragata, Isabela Island, Galapagos
Buff-tailed bumblebee (Bombus terrestris) queen about to land at her nest burrow in a garden lawn with loaded pollen sacs to provision grubs that will become future workers for her colony, Wiltshire
Oriental Magpie Robin (Copsychus saularis) female, Whitefield, Bangalore, India, March
American crocodile (Crocodylus acutus), Jardines de la Reina / Gardens of the Queen National Park, Caribbean Sea, Ciego de Avila, Cuba
Great anglehead lizard (Gonocephalus grandis) male, Tioman Island, Malaysia
Hippopotamus (Hippopotamus Amphibius) with back covered in water hyacinth on back after leaving water. Mana Pools National Park Zimbabwe
Elephant (Loxodonta africana) using termite mound to reach for food, Mana Pools National Park, Zimbabwe
Tibetan gazelle or Goa, (Procapra picticaudata) leaping, in mid air. Dulan Nature Reserve, Tibetan Plateau, Qinghai, China. October
Chital (Axis axis) female leaping. Tadoba Andhari Tiger Reserve / Tadoba National Park, Maharashtra, India
Pine grosbeak (Pinicola enucleator), perched by berries in Rowan tree, Finland, January
Wood warbler (Phylloscopus sibilatrix) amongst willow catkins, Finland, May
Common loon (Gavia immer), on a lake, Michigan, USA. June
Garibaldi (Hypsypops rubicundus) in forest of giant kelp, (Macrocystis pyrifera) off Santa Barbara Island, California, USA
Bullfinch (Pyrrhula pyrrhula) in flight, Kuusamo, Finland, March
Three juvenile Golden jackal (Canis aureus) pups running across a path, Danube Delta, Romania, July
Moose (Alces alces) bull, portrait. Grand Teton National Park, Wyoming, USA. September
Carters semaphore gecko (Pristurus carteri) standing on rocky ground. Oman, June
Puma (Puma concolor puma), young male crossing road in front of car. Estancia Amarga, near Torres del Paine National Park, Patagonia, Chile. December 2018
Yacare Caiman (Caiman yacare) gaping to regulate its body temperature, with attendant hoverflies, the Paraguay River, Taiama Reserve, western Pantanal, Brazil, South America
Ezo red fox (Vulpes vulpes schrencki) sitting on snow, licking lips after eating Vole. Hokkaido, Japan. April
Eurasian red squirrel (Sciurus vulgaris orientis) sitting on branch with nut in mouth. Hokkaido, Japan
African penguin (Spheniscus demersus) standing on rock at dusk, with clustered penguins in background. South Africa. November
Humpback whale (Megaptera novaeangliae australis) female, stationary with calf heading to surface for breath of air. Vava'u, Tonga, South Pacific
Greater flamingo (Phoenicopterus roseus) pair at rest in water, Cape Town, South Africa, July
Atlantic puffin (Fratercula arctica) swimming on the ocean surface, Svalbard, Norway, June
Grey peacock-pheasant (Polyplectron bicalcaratum) Tongbiguan Nature Reserve, Dehong, Yunnan, China
Temmincks tragopan pheasant (Tragopan temminckii) Chongqing region, Sichuan, China
Kalij pheasant (Lophura leucomelanos) Tongbiguan Nature Reserve, Dehong, Yunnan, China
Cuttlefish (Sepia officinalis) feeding on Scorpionfish (Scorpaenidae). Tenerife, Canary Islands
Blue shark (Prionace glauca). Canary Islands. North Atlantic Ocean
RF - Guanaco (Lama guanicoe) standing in front of mountain towers of Paine. Torres del Paine National Park, Patagonia, Chile. December 2018
American bison (Bison bison) female grazing near thermal pool, snow on ground. Biscuit Geyser Basin, Yellowstone National Park, USA. February 2019
Johnston's three-horned chameleon, (Trioceros johnstoni), male on tree branch, Kahuzi-Biega NP, Democratic Republic of Congo, November. Non-ex
Indian leopard (Panthera pardus fusca) on rock with forested hills beyond. Nilgiri Biosphere Reserve, India. Camera trap image. 2019
Bengal tiger (Panthera tigris) submerged in water. Ranthambore National Park, India
False gharial (Tomistoma schlegelii) resting on rock in Mpassa river. Bateke Plateau National Park, Gabon
Springtail (Dicyrtomina ornata) on oak tree leaf, Niedersechsische Elbtalaue Biosphere Reserve, Elbe Valley, Lower Saxony, Germany, December
Wild cat (Felis sylvestris) in snow, Vosges, France, February
Trumpeter hornbill (Bycanistes bucinator) feeding, captive
Collared peccary (Pecari tajacu) standing on the rainforest floor. Madre de Dios, Peru. March
Golden-collared Tanager (Iridosornis jelskii), in the Peruvian cloud forest. Wayqecha Biological Research Station, Peru. April. Cropped
Bull Skink (Liopholis multiscutata) in coastal dunes at Browns Beach on Kangaroo Island, South Australia, Autumn
RF - Basking shark (Ceterhinus maximus) feeding on plankton in Senen Cove, Cornwall, UK. June. (This image may be licensed either as rights managed or royalty free.)
Indian peafowl or blue peafowl (Pavo cristatus) Keoladeo Ghana National Park, Bharatpur, Rajasthan, India
Chamois (Rupicapra rupicapra) Ballons des Vosges Regional Natural Park, Vosges Mountains, France, October
Alpine ibex (Capra ibex) on a mountain side in autumn with European larch tree (Larix decidua) trees in the back the Alps, Gran Paradiso National Park, Italy. November
Eurasian red squirrel (Sciurus vulgaris) reflected in water, Cairngorms National Park, Scotland, United Kingdom, October
Lapwing (Vanellus vanellus) Titchwell RSPB Reserve, Norfolk, England, UK. November