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Gills of Honey Waxcap (Hygrocybe coccinea). The National Forest, Leicestershire, UK, October
Red squirrel (Sciurus vulgaris) foraging in forest, climbing down tree trunk, Cairngorms National Park, Scotland, October
Sooty albatross (Phoebetria fusca) nesting amongst Blechnum palmiforme tree ferns. Gough Island, Gough and Inaccessible Islands UNESCO World Heritage Site, South Atlantic
Atlantic yellow-nosed albatross (Thalassarche chlororhynchos) courtship display sequence. Gough Island, Gough and Inaccessible Islands UNESCO World Heritage Site, South Atlantic
Bog cranberry (Vaccinium oxycoccos) Peak District National Park, Derbyshire, England, UK, September
Bengal tiger (Panthera tigris tigris) dominant male (T29) and resident female (T27) cooling off in a watering hole Kanha National Park, Central India. Camera trap image
RF - Black Grouse (Tetrao tetrix) male peforming flutter jump display on lek, Cairngorms National Park, Scotland, UK.May (This image may be licensed either as rights managed or royalty free.)
Nilgiri marten (Martes gwatkinsii) walking on a fallen branch in Mukurthi National Park, Tamil Nadu, India
Rhesus macaque (Macaca mulatta), juveniles playing in tree, Keoladeo NP, Bharatpur, India
Darter ( Anhinga melanogaster) drying wings, Keoladeo NP, Bharatpur, India
White rhinoceros (Ceratotherium simum) male and female, face to face, Mosi Oa Tunya National Park, Zambia
White rhinoceros (Ceratotherium simum) with a long horn, Marakele National Park, South Africa
Olive-backed sunbird (Nectarinia jugularis) female returning to her nest with nesting material, Cape Hillsborough National Park, Queensland, Australia, September
Long exposure of a Dartmoor stream, Devon, England, UK, August
Lesser spotted catshark / Dogfish shark (Scyliorhinus canicula) amongst seaweed on a maerl bed, South Arran Marine Protected Area, Isle of Arran, Scotland, UK, August
View towards Higger Tor from Over Owler Tor with heather in full bloom. Abandoned millstone can be seen in the foreground. Peak District National Park, Derbyshire, UK. August 2015
Brimstone butterfly (Gonepteryx rhamni) on Devil s-bit Scabious (Succisa pratensis). Peak District National Park, Derbyshire, UK. August
Harebells (Campanula rotundifolia) growing in disused limestone quarry. Peak District National Park, Derbyshire, UK. August
The Pinnacle Stone, Curbar Edge surrounded by flowering heather. Peak District National Park, Derbyshire, UK. August
High altitude Boeri Lake in old volcano crater, Morne Trois Pitons National Park, Dominica, West Indies, Windward Islands
Old trees in the Syndicate Forest National Park, Dominica, Eastern Caribbean. February 2017
Alpine Ibex (Capra ibex) in its landscape, Valsavarenche, Gran Paradiso national park, Aosta Valley, Italy, May
Red fox (Vulpes vulpes) in deep snow emerging from den, Valsavarenche valley, Gran Paradiso national Park, Aosta Valley, Alps, Italy, December
Alpine marmot (Marmota marmota) carrying grass and other nesting material across snow to its nest while another one is on the lookout, Gran Paradiso National Park, Aosta Valley, Italy, April
Delacouras langurs (Trachypithecus delacouri) restin, Van Long Nature Reserve, Ninh Binh, VietnamDelacours langurs (Trachypithecus delacouri) restin, Van Long Nature Reserve, Ninh Binh, Vietnam. Critically endangered
Red shanked douc langur (Pygathrix nemaeus) Son Tra Nature Reserve, Vietnam
Chimpanzee female (Pan troglodytes schweinfurthii) sleeping in a nest built in a tree, Kibale National Park, Uganda, January
Female Uganda kob (Kobus kob thomasi) suckling calf. Murchisson Falls National Park, Uganda. February
Lioness (Panthera leo), aggressively rebuffing male lion banks of the Luangwa river, South Luangwa National Park, Zambia
Coalition of two African male lions (Panthera leo) resting on the banks of the Luangwa river, dry season, South Luangwa National Park, Zambia
Hawthorn tree and granite outcrop at Holwell tor, sunrise and mist, Dartmoor National Park, Devon, UK. October 2018
Marsh fritillary butterfly (Euphydrayas aurinia) feeding on Meadow thistle flower (Cirsium helenoides), Dunsdon Nature Reserve, near Holsworthy, Devon, UK. June 18
Great Mis Tor covered in snow, Dartmoor National Park, Devon, England, UK, January 2015
Great Staple Tor at dusk, Dartmoor National Park, Devon, England, UK, January
Landscape view of Great Staple Tor, Dartmoor National Park, Devon, England, UK, January 2012
American avocets (Recurvirostra americana) two during aggressive (territorial) encounter, Bear River Migratory Bird Refuge, Utah, USA, May
White pelicans (Pelecanus erythrorhynchos) at Bear River Migratory Bird Refuge with an approaching storm in May, Utah, USA, May
Yellow-headed Blackbird (Xanthocephalus xanthocephalus) male perched in marsh vegetation (Phragmites) in foreground. Bear River Migratory Bird Refuge, view toward east
Loggerhead shrike (Lanius ludovicianus) on an ice-covered fence after an ice storm, Anahuac National Wildlife Refuge, Texas, USA. January
Short-eared Owl (Asio flammeus) shaking after preening while perched on fencepost, Boxelder County, Utah, USA, May
European otter (Lutra lutra) group with one shaking off water, Kiskunsagi National Park, Hungary, October
European otter (Lutra lutra) shaking off water, Kiskunsagi National Park, Hungary, January
Forested island on Loch Lomond, Loch Lomond and the Trossachs National Park, Scotland, UK
Rainforest and river outside the Batak village of Sitio Manggapin in Cleopatra sRainforest and river outside the Batak village of Sitio Manggapin in Cleopatras Needle Critical Habitat, Palawan, Philippines
Mangroves on Tupeti Island, Southern Lagoon, Forgotten Coast, Lagoons of New Caledonia: Reef Diversity and Associated Ecosystems UNESCO World Heritage Site. New Caledonia
Coral formations at low tide in Prony Bay in the Southern Lagoon, Lagoons of New Caledonia: Reef Diversity and Associated Ecosystems UNESCO World Heritage Site. New Caledonia
Rock flagtail (Kuhlia rupestris) fish in river on the Southern Lagoon, Forgotten Coast, Lagoons of New Caledonia: Reef Diversity and Associated Ecosystems UNESCO World Heritage Site. New Caledonia
Large-eared pika (Ochotona macrotis) Serxu County, Garze Prefecture, Sichuan Province, China