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River through Virgin Komi Forests UNESCO World Heritage site, the largest virgin forests in Europe. Ural Mountains, Komi Republic, Russia. August 2016
Meerkats (Suricata suricatta), Kgalagadi Transfrontier Park, Northern Cape, South Africa, January
Young ground squirrels (Xerus inauris) fighting, Kgalagadi Transfrontier Park, Northern Cape, South Africa, January
Eastern Ring-tailed Vontsira (Galidia elegans elegans) (endemic family Eupleridae) foraging on forest floor. Marojejy National Park, north east Madagascar
Kori bustard (Ardeotis kori) male courtship display, Kgalagadi Transfrontier Park, Northern Cape, South Africa
Misty morning view of Exmoor countryside from Winsford Hill, Exmoor National Park, Somerset, UK. October 2016
Freshwater isopods (Acanthogammarus victorii) feeding on seabed, Lake Baikal, Siberia, Russia
Young Stone sculpin (Paracottus knerii), on endemic sponge (Lubomirskia baicalensis), Lake Baikal, Siberia, Russia
Clarion Lizard (Urosaurus clarionensis) displaying, IUCN Vulnerable, Clarion Island, Revillagigedo Archipelago Biosphere Reserve / Archipielago de Revillagigedo UNESCO Natural World Heritage Site
Indian wild ass (Equus hemionus khur), group standing together, Little Rann of Kutch, Gujarat, India
Sea cucumber / Sea apple (Pseudocolochirus violaceus) Rinca, Komodo National Park, Indonesia
Tropical Flounder (Bothus mancus), Socorro Island, Revillagigedo Archipelago Biosphere Reserve / Archipielago de Revillagigedo UNESCO Natural World Heritage Site (Socorro Islands), Pacific Ocean
Pristine tropical rainforest and granite cliffs in mist, Morne Seychelles National Park, Mahe Island, Republic of Seychelles
Galapagos Shark (Charcharhinus galapagensis), Roca Partida islet, Revillagigedo Archipelago Biosphere Reserve / Archipielago de Revillagigedo UNESCO Natural World Heritage Site (Socorro Islands)
Brazilian tapir (Tapirus terrestris) portrait, Tambopata Research Centre, Peru
Nurse sharks (Ginglymostoma cirratum) over seagrass at night. White spiracle behind eye is visible. South Bimini, Bahamas. The Bahamas National Shark Sanctuary. Gulf Stream, West Atlantic Ocean
Sol de Manana geyser bassin, Reserva Eduardo Avaroa, Altiplano, Bolivia
Puna / James flamingo (Phoenicoparrus jamesi) flock on Laguna Colorado, Reserva Eduardo Avaroa, Altiplano, Bolivia
Egyptian goose (Alopochen aegyptiaca) pair amongst Devil s-thorn yellow flowers (Tribulus terrestris) Etosha National Park, Namibia, March
Scimitar-horned oryx (Oryx dammah) captive in enclosure of Souss Massa National Park, Morocco. Extinct in the Wild
Hippopotamus (Hippopotamus Amphibius) territorial aggressive behavior, Chobe National Park, Botswana
Sea apple (Pseudocolochirus violaceus) Komodo National Park, Indonesia
Milky Way rising over the south wall of the Grand Canyon, Arizona, USA. November 1 2015. Composite image
European Chubb (Squalius / Leuciscus cephalus) in cave, Plitvice Lakes National Park, Croatia. November
Puma (Puma concolor) in high altitude habitat of Torres del Paine National Park, Chile
Toque Macaque (Macaca sinica) baby peering through mothers legs, Yala National Park, Southern Province, Sri Lanka
Strangler fig (Ficus sp) growing over Hoop Pine tree(Araucaria cunninghamii) with branches knotting round each other, Morans Falls Track, Green Mountains
Sumatran orangutan (Pongo abelii) female next to infant, Gunung Leuser National Park, Indonesia, UNESCO World Heritage site, November
Sumatran orangutan (Pongo abelii) female moving between lianas, Gunung Leuser National Park, UNESCO World Heritage site, November
African elephant (Loxodonta africana) feeding on grasses, Chobe River, Chobe National Park, Botswana. May
Band of wild Rapa Nui horses / colts, walking near Easter island heads on Ahu TongarikiBand of wild Rapa Nui horses/colts, walking near Easter island heads on Ahu Tongariki, Rapa Nui National Park UNESCO World Heritage Site, Easter Island / Rapa Nui, Chile
Black korhaan (Eupodotis afra) male displaying and calling. Etosha National Park, Namibia, March
African buffalo (Syncerus caffer) feeding on water lillies in Chobe River, Chobe National Park, Botswana. May
Indian rhinoceros (Rhinoceros unicornis) portrait. Manas National Park UNESCO World Heritage Site, Assam, India
Banyan tree (Ficus macrophylla) Valley of the Shadows, Lord Howe island, Lord Howe Island Group UNESCO Natural World Heritage Site, New South Wales, Australia, October 2012
Dingo (Canis lupus dingo) on the beach at night, Fraser Island UNESCO World Heritage Site. Queensland, Australia, November
Birch tree (Betula) growing in Llyn Padarn, Llanberis, Snowdonia National Park, North Wales, UK, March
Indian hog deer (Hyelaphus porcinus) mother and fawn, Manas National Park UNESCO World Heritage Site, Assam, India
Blackbuck (Antelope cervicapra), male, Tal Chhapar Wildlife Sanctuary, Rajasthan, India
Lowland Streaked Tenrec (Hemicentetes semispinosus) active on forest floor at night. Masoala National Park, north east Madagascar
Horned adder (Bitis caudalis) camouflaged in its environment, Namib Naukluft National Park, Namibia June
Peringueys / sidewinding adder (Bitis peringueyi) hidden in the sand, Naukluft National Park, Namibia
Red panda (Ailurus fulgens) family, adult female and two subadult juveniles, rest together in the canopy of the cloud forest. Singalila National Park, West Bengal, India
Polar bear (Ursus maritimus) sow with two cubs walking past a pile of Bowhead whale (Balaena mysticetus) bones, Kaktovik, Barter Island, 1002 Area, Arctic National Wildlife Refuge, North Slope
Polar bear (Ursus maritimus) spring cub sitting upright on hind legs balancing, Bernard Spit, 1002 Area, Arctic National Wildlife Refuge, North Slope, Alaska, USA, October. Vulnerable species
Scuba diver exploring volcanic crack Nesgja, in Asbyrgi National Park, northern Iceland
Nile crocodile (Crocodylus niloticus). Zimanga Private Game Reserve, KwaZulu-Natal, South Africa. September
Aldabra Giant Tortoises (Aldabrachelys gigantea) resting in a pool to keep cool, Grand Terre, Natural World Heritage Site, Aldabra
